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searchable PDF - Association for Mexican Cave Studies

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Eric Remington, William Rhodes, Jim Rodemaker,Francis Rose, J. Mark Rowland, Carol Russell, PhillipRussell, Fred Sawin, Terry Sayther, Michael Shawcross,Jim Shepperd, Rex Shepperd, Tommy Shifflett,Mark Shumate, A. Richard Smith, Jim Smith,Richard M. Smith, Carmen Soileau, Reynaldo Solis,Bill Steele, Bill Stone, Peter Strickland, Pierre Strinati,Joe Sumbera, David Sustare, Ron Syme, MillsTandy, Virginia Tipton, William Tozer, Cindy Tracy,Gary Tucker, Merydith Turner, 1. Turpin, GregWalker, Mike Wharton, Jack White, Lisa Wilk, DanWitter, and Steve Zeman. To those whose names havebeen inadvertently omitted I offer my apologies.Numerous systematists have assisted with identificationand description of the material collected fromthe caves of this area. I thank all of the following <strong>for</strong>their kindness in supplying this and much other in<strong>for</strong>mationon the cave and related epigean fauna: R. K.Allen, mayflies; D. M. Anderson, beetles; E. W.Baker, mites; T. C. Barr, Jr., beetles; S. W. T. Batra,hymenopterans; R. W. Baumann, embiopterans, mayflies,stoneflies; T. E. Bowman, mysids, isopods; A.Brindle, earwigs; H. R. Burke, beetles; G. W. Byers,flies; M. J. Campbell, beetles: R. W. Carlson, hymenopterans;O. 1. Cartwright, beetles; N. B. Causey,millipeds; F. E. Chace, shrimps; E. A. Chapin, beetles;K. Christiansen, collembolans; A. C. Cole, ants; D. R.Davis, moths; Denis Delorme, ostracods; W. E. Duellman,frogs, salamanders, lizards; Terry Erwin, beetles;R. G. Fennah, homopterans; D. C. Ferguson, moths;O. S. Flint, mayflies, dragonflies, neuropterans, trichopterans;O. F. Francke, scorpions; R. C. Froeschner,hemipterans; R. J. Gagne, flies; G. E. Gates,earthworms; W. J. Gertsch, sCOTpions, spiders, amblypygids,schizomids, ricinuleids; R. Gonzalez R.,diplurans; C. J. and M. 1. Goodnight, opilionids;A. B. Gurney, earwigs, roaches; T. F. Halstead, hemipterans;1. H. Herman, beetles; .J. 1. Herring,hemipterans; H. H. Hobbs, ]r., crayfish, shrimps, andentocytherid ostracods; H. H. Hobbs, III, shrimps,entocytherid ostracods; J. R. Holsinger, amphipods;H. F. Howden, beetles; T. H. Hubbell, crickets; C.Hubbs, fish; Leslie Hubricht, snails; P. D. Hurd, Jr.,bees; W. 1. Jellison, fleas; M. Kawakatsu, planarians;J. E. Keirans, ticks; J. M. Kingsolver, beetles; 1. V.Knutson, flies; G. M. Kohls, ticks; J. P. Kramer,homopterans; R. E. Lewis, fleas; J. D. Lynch, frogs;T. C. Maa, flies; P. M. Marsh, hymenopterans; A. S.Menke, hymenopterans; R. W. Mitchell, planarians,scorpions, fish; E. 1. Mock<strong>for</strong>d, psocids; W. B. Muchmore,pseudoscorpions; C. Mullinex, amblypygids;C. Nelson, lizards; R. Newcomer, salamanders; S. B.Peck, beetles; W. 1. Peters, mayflies; G. B. Rabb,salamanders, D. A. Rossman, snakes, lizards; J. M.Rowland, amblypygids, schizomids; C. W. Sabrosky,flies; G. A. Schultz, isopods; W. A. Shear, millipeds;A. E. Smalley, crabs; R. 1. Smiley, mites; D. R.Smith, termites, ants; M. Soleglad, scorpions; A.Solem, snails; P. J. Spangler, beetles; T. J. Spilman,beetles; G. C. Steyskal, flies; M. B. Stoetzel, homopterans;A. Stone, flies; R. W. Strandtmann, mites;W. R. Suter, beetles; S. Szerlip, hemipterans; M.Tandy, fish; F. G. Thompson, snails; W. Voss, mites;F. W. Wagner, scorpions; D. Wake, salamanders; L. M.Walkley, hymenopterans; R. E. Warner, beetles; A.Weaver, centipeds; J. P. Webb, mites; D. M. Weisman,moths; R. 1. Wenzel, beetles, flies; R. White, beetles;W. W. Wirth, flies; D. 1. Wray, collembolans; P.Wygodzinsky, diplurans, thysanurans; H. C. Yeatman,copepods; C. E. Yunker, mites.I thank the following <strong>for</strong> their reading of all orpart of the manuscript and <strong>for</strong> making many usefulsuggestions: T. C. Barr, Jr., T. E. Bowman, N. B.Causey, K. Christiansen, W. R. Elliott, F. W. Fisk,O. F. Francke, R. C. Froeschner, G. E. Gates, W. J.Gertsch, C. J. and M. 1. Goodnight, H. H. Hobbs, Jr.,H. H. Hobbs, III, J. R. Holsinger, 1. Hubricht, E. 1.Mock<strong>for</strong>d, W. B. Muchmore, S. B. Peck, J. M. Rowland,W. H. Russell, G. A. Schultz, W. A. Shear, A. R.Smith, H. M. Smith, T. J. Spilman, j. Sullivan, F. W.Wagner, A. Weaver, P. Wygodzinsky, and H. C. Yeatman.I take full responsibility <strong>for</strong> all errors and <strong>for</strong>all interpretations of the data.Jerry Atkinson, David McKenzie, William H.Russell, and Peter Sprouse assisted in the preparationof the locality list. William R. Elliott and Hal Storyprovided invaluable aid in the preparation of themaps.Financial support <strong>for</strong> some of the field work inMexico was provided by the National GeographicSociety and The Museum, Texas Tech University.Additional assistance was provided by Nell B. Causey,Willis J. Gertsch, and Theodore H. Hubbell.HISTORYMexicoThe first biological investigation of a cave inMexico was conducted by Dominik Bilimek on January14, 1866, when he visited Grutas de Cacahuamilpa,Guerrero (Bilimek, 1867). Among the II speciesreported was one troglobite, Lepisma anophthalma.The next important contribution to ourknowledge of the cavernicole fauna of Mexico wasthat of the distinguished <strong>Mexican</strong> zoologist, D. A. 1.Herrera (1891). Among species he collected in Grutasde Cacahuamilpa was the same species of troglobite,described as Lepisma cacahuamilpensis. Herrera also5

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