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*Gruta Sur de San Bartolo, 9 km S Santa Catarina,900 m, Mpo. de Santa Catarina (Raines, 1968b:140-141,143) (Garza Garcia, G14C25)Cueva de Tio Bartolo-see Gruta Norte de San Bartoloand Gruta Sur de San BartoloCanon de Iturbide Region+*Cueva La Charrera, 27.3 km SW Linares, Mpo. deLinaresCueva del Ehanito-see Cueva del Nacimiento de ElEhanito+Cueva de Guadalupe, Canon de Itiurbide, 25 km SWLinares, Mpo.de Linares (Malaga Alba and Villa R.,1957:539)*Cueva del Nacimiento de El Ebanito, El Ebanito,10 km ENE Iturbide, Mpo. de Linares (Russell,1973b:280) (lturbide, G14C67)Cerro de la Boca Region*Cueva de Chorros de Agua, 23 km SW Montemorelos,700 m, Mpo. de Montemorelos (Russell and Raines,1967:29,38) (Montemorelos, G14C47)Cerro Potosi Region+Cave, Lorna La Cueva, 1.5 km SW San Jose La Boya,27 km WNW Galeana, 2720 m, Mpo. de Galeana(Radovsky et a!., 1971:738) (Galcana, G14C56)Cuesta de Chipinque RegionSmall caves on Cuesta de Chipinque-see Caves 1-3*Cave No.1, Cuesta de Chipinque, Monterrey, 1500 m,Mpo. de Monterrey (Peck and Peck, 1973:64)(Garza Garcia, G14C25)Cave No.2, Cuesta de Chipinque, Monterrey, 1500 m,Mpo. de Monterrey (Peck and Peck, 1973:64)(Garza Garcia, G14C25)Cave No.3, Cuesta de Chipinque, Monterrey, 1500 m,Mpo. de Monterrey (Peck and Peck, 1973:64)(Garza Garcia, G14C25)Galeana RegionChevy Sink, 7 km SW Galeana, 1780 m, Mpo. deGaleana (Galeana, G14C56)Pozo de Gavilan, 7 km SW Galeana, 1780 m, Mpo. deGaleana (Russell and Raines, 1967:31, 38, 40m)(Galeana, G14C56)Cueva del Limon, 1.2 km E San Marcos, 15 km SGaleana, Mpo. de Galeana (San Jose de Raices,G14C66)*Resumidero de Pablillo, 4 km NNE Pahlillo, 26 kmSE Galeana, 2000 m, Mpo. de Galeana (Russelland Raines, 1967:31,38, 41m) (Iturhide, G14C67)Sotano de la Pina-see Sotano del PinoSotano del Pino, 6 km SSW Galeana, 1700 m, Mpo.de Galeana (San Jose de Raices, G14C66)Cueva Sur del SOlano de Dos Cuevas, 1.5 km S SantaFe, 15.5 km ESE Galeana, 1800 m, Mpo. de Galeana(Reddell, 1966d:49) (San Jose de Raices,G14C66)Cueva de Yeso Blanco, 6 km SW Galeana, 1740 m,Mpo. de Galeana (Galeana, G14C56)Monterrey Region+Well, Monterrey, Mpo. de Monterrey (Packard, 1894:732)Pico de Carrizal Region*Gruta de Carrizal, 10 km SW El Candela, Mpo. deLampazos de Naranjo (Russell and Raines, 1967:13,14m)Potrero Redondo RegionSotano de la Anticlina, 9 km SSW Villa de Santiago,Mpo. de Villa de Santiago (Raines, 1972b:96)(Allende, G14C36)Sotano de Potrero Redondo, 17 km S Villa de Santiago,Mpo. de Villa de Santiago (Allende,G14C36)Redondo Pit Cave-see Sotano de Potrero RedondoPurificacion RegionCueva del Borrego-see TAMAULIP ASCueva del Brinco--see Sistema Purificacion, TAMAlJ-LIPASCueva de California-see TAMAULIPASCueva del Camino-see TAMAULIPASCueva de Chuparosa-see TAMAULIPASSotano de.Jesus-see TAM AU LIP ASCueva del Ojo de Agua, 4 km WSW Dulces Nomhres,Mpo. de Aramherri (Casas Reales, Fl4A18)*Cueva Chica del Ojo de Agua, 4 km WSW DulcesNombres, Mpo. de Aramherri (Casas Reales,Fl4A18)Cueva de los Parajitos, 2 km N Tinajas (=Los Tanques),7 km SE Dulces Nombres, Mpo. de Aramberri(Casas Reales, F14A18)Pozo de los Penuelos-see Sotano de las Penue!as*Sotano de las Penue!as, 3 km SW Dulces Nombrcs,2010 m, Mpo. de Aramberri (Casas Reales,F14A18)Sotano de Rancho Nuevo n. I-see TAMAULIPASSotano de Rancho Nuevo n. 2-see TAMA ULIP ASSotano de Rancho Nuevo n. 3-see TAMAULIPASSotano de Rancho Nuevo n. 4-see TAMAULIPASCueva de Rancho Revilla-see Cueva de Revilla,TAMAULIPASCueva del Tecolote-see Cueva de \)esmontes, TAMAULIPASCueva Vrincon-see Sistema Purificacion, TAMA ULIPASSierra Las Animas Region+Puente de Dios, 7 km NNW Galeana, 1460 m, Mpo.de Galeana (Russell and Raines, 1967: 38) (Galeana,G14C56)Sierra del Fraile RegionGrutas de Garcia-see Grutas de Villa de Garcia+Cueva del Rincon de la Virgen, 6.8 km N Villa deGarcia, 1200 m, Mpo. de Villa de Garcia (MalagaAlha and Villa R., 1957:559,564,566) (Hidalgo,G14C15),x'Grutas de Villa de Garcia, 7 km NW Villa de Garcia,1060 m, Mpo. de Villa de Garcia (Russell andRaines, 1967:19, 20m)(Hidalgo, G14C15)Sierra de Garia RegionCueva de Constantin, 8 km S Espinazo, Mpo. deMina (Bittinger, 1972:56, 57m) (Reata, G14A83)Sierra de Gomas RegionCave 7 km SSW Bustamante, Mpo. de Bustamante298
Pit 7 km SSW Bustamante, Mpo. de Bustamante(Fromen, 1965:44-45 )*Grutas del Palmito, 7 km SSW Bustamante, Mpo. deBustamante (Russell and Raines, 1967:16, 1718m; Ediger, 1970:3-7, 5m)Cueva del Precipicio, 9 km SW Bustamante, Mpo. deBustamante (Russell, 1973a:363-365; Walsh, 1973:100-101)Sierra de Iguana Region+Cueva del Diablo, 7 km by road W Sabinas Hidalgo,Mpo. de Sabinas Hidalgo (Hershberger, 1967:8889,90m)+Cueva Envenenada, 7 km by road W Sabinas Hidalgo,Mpo. de Sabinas Hidalgo (Aguirre Pequeno, 1959:245,246)Sierra de Santa Clara RegionCueva del Camino-see Cueva de las FisurasCueva de las Fisuras, 16 km by road W Sabinas Hidalgo,Mpo. de ?Sabinas Hidalgo (Russell and Raines,1967:21)Sierra de la Silla Region*Flooded mine below Cueva de la Boca, 3 km E Villade Santiago, 500 m, Mpo. de Villa de Santiago(Peck and Peck, 1973:63) (Allende, G14C36)*Cueva de la Boca, 3 km E Villa de Santiago, 540 m,Mpo. de Villa de Santiago (Russell and Raines,1967:26,35, 36m, 37) (Allende, G14C36)Cueva de la Villa de Santiago-see Cueva de la BocaSierra EI Tarillal RegionCueva de las Vigas, EI Carmen (=Las Vigas), 22 km EArteaga, 2300 m, Mpo. de Santa Catarina (Reddell,1966d:53) (San Antonio de las Alazanas, G14C35)Zaragoza RegionOAXACACueva de Cuesta Blanca, 3 km W Zaragoza, Mpo. deZaragoza (Russell and Raines, 1967: 31, 38) (Zaragoza,Fl4A17)Sotano de Cuesta Blanca, 3 km W Zaragoza, Mpo. deZaragoza (Russell and Raines, 1967:38) (Zaragoza,Fl4A17)Dolina Cave, 4 km N Zaragoza, Mpo. de Zaragoza(Zaragoza, F14A17)Acatlan Region+Cave on Cerro Pinon, 12 km S Acathin, Mpo. deAcatlan (Constantine, 1966:126)Cave near Los Corrales, 11 km SSW Acatlan, Mpo. deAcatlan+Cave 1.5 km NW Los Limones, 18 km S Acatlan,Mpo. de Acatlan (Constantine, 1966:125)+Cave 5 km W Los Limones, 18 km S Acatlan, Mpo.de Acatlan (Constantine, 1966:125)Cueva de Campo Chico, 9 km SSW Acatlan, Mpo. deAcatlan*Cueva de Cayetano, 10 km SW Acatlan, Mpo. deAcatlan*Cueva de Los Corrales, 2 km W Los Corrales, 11 kmSSW Acatlan, Mpo. de Acatlan*Cueva de la Culebra, 10 km SW Acatlan, Mpo. deAcatlan*Cueva Desapareciendo, 2 km SW Acatlan, Mpo. deAcatIanSotano Desapareciendo, 2 km SW AcatJan, Mpo. deAcatlan*Cueva de la Finca, 10 km SW Acatlan, Mpo. de AcatIan*Cueva de Juan Sanchez, 10 km NW Acatlan, Mpo. deAcatlan (Byrd, 1976:23, 24m)*Cueva Chica de Juan Sanchez, 12 km NW Acatlan,Mpo. de AcatlanCueva de la Junta, 5 km SW Acatlan, Mpo. de AcatIan (Reddell, 1973e:88-S9)*Cueva de la Laguna Verde, 10 km SW Acatlan, Mpo.de Acatlan (Coons, 1977:18-20, 21m)*Cueva del Lencho Virgen, 9 km SSW Acatlan, Mpo.de Acatlan (Reddell and Elliott, 1974:9)*Cueva de las Maravillas, 6 km SSW Acatlan, Mpo. deAcatJan (Reddell, 1973f:95)+Cueva del Nacimiento, above Vista Hermosa, Acatlan,Mpo. de Acatlan (Villa R., 1967:224)*Cueva del Nacimiento del Rio San Antonio, 10 kmSSW Acatlan, Mpo. de Acatlan (Reddell, 1973e:89; Reddell, 1973f:95; Reddell and Elliott, 1974:9)Cueva Sin Fin, 2 km SW Acatlan, Mpo. de AcatlanSotano Sin Hondo, 5 km SSW Acatlan, Mpo. deAcatlanHuautla de Jimenez Region*Cave near Puente de Fierro, 1.5 km N Huautla deJimenez, Mpo. de Huautla de JimenezSink near Huautla de Jimenez, Mpo. de Huautla deJimenezCave No.3, near Huautla de Jimenez, Mpo. deHuautla de JimenezPit No.2, near Huautla de Jimenez, Mpo. de Huautlade Jimenez*Sotano de Agua de Carrizo, 5 km ESE Huautla deJimenez, Mpo. de Huautla de Jimenez (Stone,1979a:33-52,2 maps)Cueva del Arroyo Mazateca, San Miguel Dolina, SanMiguel, 5 km SE Huautla de Jimenez, Mpo. deHuautla de JimenezCueva del Camino, Puente de Fierro, 1.5 km NHuautla de Jimenez, Mpo. de Huautla de Jimenez*Cueva del Escorpion, San Miguel Dolina, San Miguel,5 km SE Huautla de Jimenez, Mpo. de Huautla deJimenezThe Fissure-see La Grieta*La Grieta, 2 km E San Andres, 5 km E Huautla deJimenez, Mpo. de Huautla de Jimenez (Stone,1977:7-17, map; Steele, 1978:26-29, map)*Milliped Cave, 5 km SE Huautla de Jimenez, Mpo. deHuautla de Jimenez (Fish and Russell, 1966:63,66m)*Milliped Cave, 1.5 km N Huautla de Jimenez, Mpo.de Huautla de Jimenez (Russell, 1965 :62)*Cueva del Molino de Carne, 5 km ESE Huautla deJimenez, Mpo. de Huautla de Jimenez*Cueva de los Pajaros, 5 km SE Huautla de Jimenez,Mpo. de Huautla de JimenezCueva Arriba del Presidente, 1 km N Huautla deJimenez, Mpo. de Huautla de Jimenez*Cueva Bonita del Presidente, 1 km N Huautla deJimenez, Mpo. de Huautla de Jimenez299
Page 1:
A REVIEW OF THECAVERNICOLE FAUNA OF
Page 8:
. 1-----,, ,", ,\'"I/I~" )/Fig. I.-
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collected a troglobitic cirolanid i
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C. Bolivar and J. Carranza investig
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springs in the vicinity of Cautro C
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Cueva del Diablo near Orizaba, Vera
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Those species of troglobite which h
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Table 2.-(continued)1970. Eodrilus
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CAVE REGIONSOnly the cave regions o
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Table 3.-(continued)TroglobitesTrog
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y the waters of the Amistad Reservo
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Although a few caves in the Norther
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an elevation of about 900 meters. T
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the gryllid cricket Paracophus suba
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has been Jiscussed by Segerstrom (1
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only apparent troglophile is the ps
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The cave fauna of this region is ri
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igneous rock immediately east of th
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Rio Soto la Marina and terminates a
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caves studied are many large stream
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unusual species is Neogovea mexasca
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pallidus Rowland, the nesticid spid
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troglobites: the trichoniscid isopo
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Cambridge, and Pholcophora bispinos
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of the genus Lutzomyia has not been
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The principal cavernous fonnation i
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ocks of Paleocene to Pliocene age.
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Beach and Supra-tidal Zone, the I\o
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The aquatic fauna of Mexico and Cen
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oaxacae Hobbs from the Valle Nacion
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described, the genus occurs in cave
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large genus with numerous epigean s
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milliped, laniatorid harvestmen, an
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Table 4.-Summary of cave inhabiting
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Discussion.-This large, troglobitic
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any way a part of the true cave fau
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they have been washed in with soil
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part of the aquatic fauna. A very l
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The genus Helicodiscus is a signifi
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associated with true cave waters. P
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Family DaphnidaeWilson (1936) repor
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tul), Cueva de San Isidro, Cueva de
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30 m below entrance of Cueva de la
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underflow of rivers, alluvial grave
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the similarities between the distri
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Yucatan but did not identify the sp
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Protrichoniscus: Reddell, 1967a:24.
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It indudes five spccies, of which t
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Type-locality.-Pozzo, Etla, Oaxaca,
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Records.-Campeche: Volc
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164; Pearse, 1945:167, 168, fig. 13
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Family AtyidaeThe family Atyidae is
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Creaseria morleyi (Creaser)Blind sh
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the pond which is fed by the cave s
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philes. Pseudothelphusa (Pseudothel
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Typhlochactas elliotti MitchellTyph
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Distribution.-Known only from the t
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(Dumitresco and Orghidan, 1977); th
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Leucohya magnifica MuchmoreUndescri
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Discussion.-This is the largest spe
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Type-Iocality.-S6tano de Huitzmolot
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Bartolo is a name applied to two ne
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Rowland, 1975b:19, 20; Dumitresco,
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Type-Iocality.-Actlin Kaua, Yucatan
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Distribution.-Known only from caves
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een collected. It is frequently fou
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Distribution.-Known from two caves
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actually a natural cave artificiall
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Discussion.-This large species has
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and T. patellaris Bryant from caves
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estricted to the Mexican Plateau (s
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maya Chamberlin and Ivie from Campe
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Metagonia chiquita GertschMetagonia
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Metagonia capilla: Mitchell and Kaw
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the cave and in general association
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DiscWlSion.-This is a delicate, eye
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Type-Iocality.-Cueva de Los Sabinos
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family. Clarence and Marie Goodnigh
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n. 2, San Luis Potosi; and K. rugos
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Type-locality.-Cueva de Apoala, San
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Records.-Chiapas: Sumidero de Canad
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Family HeterozerconidaeSpecimens of
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only one species. All known epigean
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nation must await study, but it is
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Mexico and Guatemala, and several u
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Type-locality.-Gruta de Atoyac, Ver
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Records.-San Luis PotOSI: Sotano de
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Records.-Veracruz: Grutas de Atoyac
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the genus Synoptura and removed the
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known from caves all contain troglo
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las Perlas, Cueva Chica de la Perra
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Discussion.-This species is distinc
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Speodesmus pecki ShearPolydesmidae,
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appears to be most closely related
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this is almost certainly M. russell
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Table 21.-Summary of cave inhabitin
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Type-Iocality.-Cueva Bonita del Pre
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Type-Iocality.-Cueva de El Jobo, El
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Order DipluraFamily CampodeidaeAlth
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of small soil inhabiting japygoids
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(Linnaeus) has been collected from
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Cave. Paracophus caecus appears to
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the most poorly known species in th
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Family EpipsocidaeThe family Epipso
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citata Van Tyne in Cenote de Piste,
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een found in caves and mines in Yuc
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ground dwelling habit, many beetles
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Type-Iocality.-Sotano de Tejamanil,
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Vigna Taglianti, 1972:118, 121-127,
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Veracruz) belongs in another subgen
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Troglobacanius sbordonii VomeroTrog
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Ptomaphagus (Adelops) is a group of
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(A.) speLaeus, from Grutas de Cacah
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epresented by many species. Tenebri
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Myodopsylla gentilis Jordan and Rot
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genus Pholeomyia have been collecte
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Wheeler (1938) has reported the pre
Page 246 and 247:
263.264, 265, 267, 270, 273, fig. 5
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Anophthyctys jordani: Stefanelli, 1
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Prietella phreatophila Carranza, 19
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TyphlilJs: Pearse, 1938a:13, 15; Hu
Page 254 and 255: Family BufonidaeFour species of toa
Page 256 and 257: Oaxaca; and E. yucatanensis Lynch f
Page 258 and 259: sity of bats in the tropics. In add
Page 260 and 261: Three species of short-tailed bats
Page 262 and 263: abundant in the entrance area of Cu
Page 264 and 265: Andrews, E. W., IV. 1970. Balankanc
Page 266 and 267: Bolivar y Pieltain, C. 1940. Explor
Page 268 and 269: Brignoli, P. M. 1979a. The morpholo
Page 270 and 271: Conde, B. 1975. Description du prem
Page 272 and 273: Ediger, G. 1970. The Birthday Passa
Page 274 and 275: Gonzalez Ochoa, A. 1963b. Relacione
Page 276 and 277: Hobbs, H. H.,Jr., and T. C. Barr, J
Page 278 and 279: Jordan, c. B. 1937. Bringing in the
Page 280 and 281: McKenzie, D. 1965a. Caves of the Si
Page 282 and 283: Monod, T. 1975. Sur la distribution
Page 284 and 285: Peck, S. B., and J. J. Lewis. 1977.
Page 286 and 287: Reddell, J. R., and W. R. Elliott.
Page 288 and 289: Sanborn, C. C. 1936. Records and me
Page 290 and 291: Stone, B. 1979. The 1979 San Agusti
Page 292 and 293: Villa R., B. 1956. Una extrana y se
Page 294 and 295: Wimsatt, W. A. 1969. Transient beha
Page 296 and 297: +*Cueva de los Chivos, 5 km E Altam
Page 298 and 299: Sumidero Yochib, Yochib, 8 km NE Te
Page 300 and 301: Serrania del Burro RegionCave, Haci
Page 302 and 303: +Cueva de Tezoapa, 1.5 km E Acahuiz
Page 306 and 307: *Cueva del Puente de Fierro, 1.5 km
Page 308: Cueva de los Riscos, 2 km S Rancho
Page 311 and 312: Cueva de El Coy-see Cueva del Nacun
Page 313 and 314: +*Cueva del Aire, 1.8 km at S85°W
Page 315 and 316: *Sistema Purificacion, Conrado Cast
Page 317 and 318: *Cueva del Nacimiento del Rio Frio,
Page 319 and 320: Sotano de Humo, 5 km NW San Andres,
Page 321 and 322: and Bohor, 1975:25-27, 26m; Reddell
Page 323 and 324: *Actun Xpukil, 3 km S Calcehtok, Mp
Page 325 and 326: Appendix 2LIST OF TROGLOBITES BY ST
Page 327 and 328: Paravachonium superbum-I'seudoscorp
Page 329 and 330: SUPPLEMENTWhile this manuscript was
Page 331 and 332: Cambala speobia (Chamberlin) (p. 19
Page 333: mida (Arachnida) in the New World.
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