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searchable PDF - Association for Mexican Cave Studies

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(Villalobos). Endemic aquatic troglobites include thediaptomid copepod Diaptomus (Microdiaptomus)cokeri Osorio Tafall, the microcerberid isopod Microcerberusn. sp., and the palaemonid shrimp Troglocubanusperezfarfanteae Villalobos. The terrestrialtroglobite fauna includes the rhagidiid mite Rhagidiaweyerensis (Packard), also known from caves in theUnited States. Other species known from otherregions include the squamiferid isopod Spherarmadillocavernicola Mulaik, the trichoniscid isopodBrackenridgia bridgesi (Van i\arne), the opilionidHoplobunus boneti (Goodnight and Goodnight), thescolopendrid centipede Newportia sabina Chamherlin,and the collembolan Pseudosinella petrustrinatiiChristiansen. Endemic troglobites are the trichoniscidisopod Cylindroniscus vallesensis Schultz, thechactid scorpion Typhlochactas elliotti !\Iitchell, thechthoniid pseudoscorpions Aphrastochthonius pan'usMuchmore and A. russelli \Iuchmore, the vachoniidpseudoscorpions Paravachonium n. sp. and P. bolivariBeier, the protoschizomid Agastoschizomus luciferRowland, the schizomids Schizomus cookei K.owlandand S. mitchelli K.owland. the amhlypygid Paraphryllusbaeops Mullinex, the pholcid spider Metagoniapachona Gertsch, the ricinllleids Cryptocellusn. sp. and C. osorioi Bolivar, the rhagidiid miteRhagidia trisetata Elliott and ~trandtrnann, the trichopetalidmilliped JIexiterpes sabinus Causey, thetrichopolydesmid milliped Speodesmus pecki Shear,an undescribed genus and species of Jlicoletiid thysanuran,the kinnarid homopteran Oeclidius hadesFennah, and the histerid bectle Troglobacanius bolivariVomero.The aquatic troglophile fauna includes several speciesof protozoans, rotifers. ostracods, copepods, thecrayfish Procambarus (Ortmannicus) aeutus cuevachicae(Hobbs), and th~~ characiu Astyanax mexicanus(Phillipi). About 100 terrestrial species arc prohabletroglophiles in the Sierra de EI Ahra. [t is not possiblehere to do more than list some of the more distinct orwidespread species. The arachnid fauna includes theschizomid Schizomus mexicanus I{owland, the amblypygidParaphrynus pococki \Iullinex. the ricinuleidCryptocellus pelaezi Coronado, the opilionidsCynorta jamesoni Goodnight and Goodnight andKaros parous (;oodnight and Goodnight, and the palpigradeKoenenia hanseni Silvestri. Troglophilicspiders include the ctenid Ctenus mitchelli Gertsch;the leptonetid Leptoneta rainesi Gertsch; the mysmenidMaymena chica Gertsch; the nesticids Eidmannellapallida (Emerton) and Eidmannella n. sp.;the pholcids Metagonia pasquinii Brignoli, Metagoniatinaja Gertsch, 1Hodisimus boneti Gertsch, Ph 01­cophora elliotti Gertsch, and Physocyclus globosus(Taczanowski); and the scytodid Loxosceles valdosaGertsch. The milliped fauna includes the pyrgodesmidsMyrmecodesmus errabundus (Shear) and M.sabinus (Chamberlin) and the spirostreptid Orthoporusmimus Chamberlin. Apterygote insect troglophilesinclude the campodeid dipluran Campodea(Campodea) chica Wygodzinsky, the japygid dipluranAllojapyx allodontus (Silvestri), amI thecollembolan Acherontiella sabina Bonet. Two speciesof troglophile gryllid cricket are known, Paracophusapterus Chopard and P. placonotus Hubbell. i\umerousspecies of beetle are tro~lophiles in the region.These include the carabid Pachyteles urrutiai Hoi Ivar,the leiodids Dissochaetus aztecus ~zymczakowskiand Ptomaphagus (£ldelops) elabra Peck, scydmaenidsof the genera "Connophron " and Euconnus(Napochus), and staphylinid beetl(~s of the generaBelonuchus, Homaeotarsus, and Stilicolina. Both thetroglophile and troglobite fauna of this region aretypical of the Sierra Madre Oriental, with its closestaffinities lying with the Sierra de Guatemala to thenorth and somewhat less with that of the ranges tothe west and sou th west.Altas Cumbres, Tamaulipas.-This is a poorlydefined region located to the southwest of CiudadVictoria on the highway to Tula. Of the three cavesknown <strong>for</strong> the region, only two havl~ been investigatedwi th respect to their invertebrate fauna. Bothare small fissure-like caves with a limited fauna. Offour invertebrates identified, th ree are troglophiles:the leptonetid spider /,eptoncta bonita Certsch, thepholeid spider Modisim us reddelli G (~rtsch, and thegryllid cricket Paracophus apterus Chopard. Thesearc typical elements of the cave fauna of the generalregIOn.Sierra de Guatemala, Tamaulipas.-The Sierra deGuatemala is a range located immediat~:ly north ofthe Rio Boqllillas and to the west of the town ofGomez Farias. It is hounded on the north by the RioGuayalejo; on the west it deseends abruptly to theplains east of JaUll1ave. Elevations range from 170 toabout 2,200 meters. Martin (1958) has summarizedthe geology, physiography, vegetation, and dirnateof the region. His study of the reptiles allll amphibiansof the region also includes mudl useful hiogeographicin<strong>for</strong>mation. The principal limestone unitis the EI Abra (= El Doctor) Limestone (Russell andRaines, 1967; Priego de Wit, 1974). Most of thedrainage is internal, with water flowing from twomajor springs, the Nacimiento del Rio Frio to thesouth and the Nacimiento del Rio Sabinas to thenorth. These springs are vauclusian in nature. <strong>Cave</strong>sare known throughout the area, but most are small.The town of Gomez Farias is built on a small ridge of36

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