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searchable PDF - Association for Mexican Cave Studies

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darkness along the cave wall. One of these is A.porteri Banks, a common troglophile in the caves ofCentral Texas. Several species of Theridion, Thymoites,and Tidarren have been occasionally collected,but their ecological status is not known.Family UIoboridaeThe genus Philoponelw is known from many cavesthroughout Mexico. Philoponella signatelw (Roewer)is an apparent troglophile in caves in Oaxaca, Puebla,and Yucatan. Philoponella semiplumosa (Simon) isknown from caves in Tamaulipas and Veracruz.Philoponelw vicina (0. P.-Cambridge) has been takenin several caves in Tabasco and Veracruz.Order RicinuleiFamily RicinoididaeThe order Ricinulei includes only one family andtwo living genera, Ricinoides in Africa and Cryptocellusin the New World. Cryptocellus ranges northfrom South America into southern Texas. In southernMexico it has been found in Chiapas, Guerrero, andYucatan; it occurs north along the Sierra Madre Orientalfrom Veracruz to Nuevo Leon, with isolatedcave populations in Durango. A single species, C.dorotheae Gertsch and Mulaik, is known from Texas.One cave species, C. cookei Gertsch, is known fromCueva Jobitzinaj, near Flores, EI Peten, Guatemala(Gertsch, 1977a). Ten <strong>Mexican</strong> species have beendescribed, of which two (c. spinotibialis Goodnightand Goodnight from Chiapas and C. gertschi Marquezand Conconi from Veracruz) a.re exclusively epigean.One species (c. pelaezi Coronado) is known fromboth cave and epigean sites in the Sierra de EI Abra,San Luis Potosi and Tamaulipas, and from epigeansites in the Sierra de Guatemala, Tamaulipas. Theremaining seven species are all known only fromcaves (see Fig. 45). Three of these are considered tobe troglobites and are listed below. The remainingfive species are either troglophiles or of uncertainecological status. An epigean species from NuevoLeon and a troglobitic species from Cueva de Valdosa,San Luis Potosi, await description.Chamberlin and Ivie (1938a) described the first<strong>Mexican</strong> cavernicole species, C. pearsei, from Grutasde Balankanche and Cueva Oxolodt, Yucatan. Thisspecies has since been found in nine additionalYucatan caves (Gertsch, 1977a). Cryptocellus bonetiwas described by Bolivar (1941) from Grutas deCacahuamilpa, Guerrero; it is also known fromGrutas de Acuitlapan, Guerrero (Coronado, 1970).Cryptocellus pewezi was described by Coronado(1970) from Cueva de Taninul n. 1, San Luis Potosi.This species has become the best known of allricin'uleids, due to the outstanding work of Dr.Robert W. Mitchell and his students at Texas TechUniversity. Pittard and Mitchell (1972) studied theextemal morphology of all life stages; Cooke (1971)studied the mating behavior; and Mitchell (1970)studied various aspects of the population dynamicsin Cueva de la Florida, Tamaulipas. Other studies,including the life cycle and the development of thethird leg, are in preparation. Gertsch (1971b) describedthree additional species of cavernicole ricinuleid.In addition to the troglobite, C. reddelli, headded C. mitchelli from Cueva del Guano, Durango,and C. bolivari from Sumidero del Camino andGrutas de Zapaluta, Chiapas. Brignoli (1974c) describedC. sbordonii from Chiapas and tentativelyreported C. bolivari from Cueva del Tio Ticho,Chiapas.Epigean ricinuleids are usually found beneathrocks along hillsides or under permanent groundcover. The cavemicolous species have been foundon guano, silt, or rarely beneath rocks. The presenceof C. pearse~ C. osorioi, and C. pewezi in vast numbershas proven that the order is not the incrediblyrare group of animals that it was once thought to be.Cryptocellus osorioi BolivarCryptocellus osorioi Bolivar, 1946:24-28, fig. 1-7;Bolivar, 1952:296; McKenzie, 1965a:36; Reddell,1967b:82; Reddell, 1967f:99; Barrera, 1968:309;Beck and Schubart, 1968:68,69,70,74; Kaestner,1968:207; Mitchell, 19691:137; Mitchell, 1969d:letter; Woolley, 1969: 151; Coronado, 1970:48,61; Mitchell, 1970b:64, 71-72, 73, 74; Pittard,1970:3,6, 10; Cooke, 1971:9, 17,18,19,20,21,22; Gertsch, 1971b:127; Reddell, 1971b:3, 34,frontispiece; Reddell and Mitchell, 1971a:148,fig. 8; Reddell and Mitchell, 197c:2; Pittard andMitchell, 1972:4, 6; Sbordoni and Argano, 1972:9, 11; Kawakatsu, 1973b:255; Reddell, 1973a:34;Reddell and Elliott, 1973a:174; Brignoli, 1974c:154, 155, 156, 157, 159,160,166,167-168,169,fig. lC; Marquez and Conconi, 1974:82, 83, fig.20; Vomero, 1974:345; Dumitresco and Juvara­Bals, 1976:163; Mitchell et al., 1977:53.Cryptocoellus osorioi: Anonymous, 1942a:221 (erroneousspelling).Cryptocellus osorici: Dumitresco and Juvara-Bals,1976: 163 (erroneous spelling).Cryptocellus ossorioi: Dumitresco and J uvara-Bals,1976:164, 178 (erroneous spelling).157

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