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Cueva de los Riscos, 2 km S Rancho Huichale, 8 kmSW Jalpan, Mpo. de J alpan (Fish and Reddell,1967:85-86,87m)Laguna Colorada RegionSinkhole, Laguna Colorada, 23 km W Xilitla, Mpo. deLanda de MatamorosCueva de Camposantos-see Sotano de CamposantosSotano de Camposantos, 5 km W El Lobo, 22 km WXilitla, Mpo. de Landa de Matamoros (Fish andReddell,1967:86)+*Cueva del Madrono, 1.5 km S El Madrono, 19 km WXilitla, 1810 m, Mpo. de Landa de Matamoros(Bonet, 1953a:92, 94-96, 93m; Sbordoni andArgano, 1972: 12-13)"Cueva del Nino,S km W El Lobo, 22 km W Xilitla,Mpo. de Landa de Matamoros (Sbordoni and Argano,1972:13)*Cueva de las Tablas, 5 km W El Lobo, 22 km WXilitla, Mpo. de Landa de MatamorosPinal de Amoles Region*Iron (7) mine, 2 km E Pinal de Amoles, Mpo. de Pinalde Amoles (Peck and Peck, 1973:67)ll5-ft. blind pit, San Gaspar, 10 km NW Pinal deAmoles, Mpo. de Pinal de AmolesCave No.1, 20 km N Pinal de Amoles, Mpo. de Pinalde Amoles (Greer, 1979:109)Cave No. 14, 20 km N Pinal de Amoles, Mpo. dePinal de Amoles (Greer, 1979:112-113, 112m)Cave No. 29-see Cave No. 14Cave No. 33-see Cave No.1*Sotano del Buque, 20 km N Pinal de Amoles, Mpo.de Pinal de Amoles (Jefferys, 1979:70-74, map)Cueva de Chevron, 3 km E Pinal de Amoles, Mpo. dePinal de Amoles (Fish and Reddell, 1967:85)Sotano de Dos Arafias Grandes, 1 km ENE RioBlanco, 2000 m, Mpo. de Pena Miller (Reddell,1979:99)(Pena Miller, F14C47)Cueva Encantada, 3 km W La Cienega, 20 km NNEPinal de Amo)es, Mpo. de Pinal de Amoles (Greer,1979:114,115m)Sotano Encantado-see Cueva Encantada"Sotano de la Escuela, Llano de San Francisco, 10 kmNW Pinal de Amoles, Mpo. de Pinal de Amoles(Bittinger, 1973:86)Sotano del Gobernador, 2 km S Pinal de Amoles,Mpo. de Pinal de Amoles (Reddell, 1979:100)"Cueva del Judio, 2 km S Pinal de Amoles, Mpo. dePinal de Amoles (Reddell, 1979:100)Sotano de La Lagunita, 2 km NE Rio Blanco,2100 m, Mpo. de Pena Miller (Reddell, 1979:99)(Pena Miller, Fl4C47)Cueva del Mercurio, Tejamanil, 3 km SW Pinal deAmoles, Mpo. de Pinal de Amoles (Reddell, 1979:100)"Cueva de los Otates, Los Pinos, 15 km NW Pinal deAmoles, Mpo. de Pinal de Amoles (Bittinger,1973:86)Sotano de la Paila, 20 km N Pinal de Amoles, Mpo.de Pinal de Amoles*Cueva de Penalta, 10 km NW Pinal de Amoles, Mpo.de Pinal de AmolesSotano del Pino, 20 km NNE Pinal de Amoles, Mpo.de Pinal de Amoles*Cueva del Puerto del Leon, 4 km E Rio Blanco,Mpo. de Pinal de Amoles (Reddell, 1979:99)(Pena Miller, Fl4C47)Cueva de Tejamanil, 0.5 km NE Tejamanil, 2.5 kmSW Pinal de Amoles, Mpo. de Pinal de Amoles(Fish and Reddell, 1967:85)*Sotano de Tejamanil, 0.5 km NE Tejamanil, 2.5 kmSW Pinal de Amoles, Mpo. de Pinal de Amoles(Fish and Reddell, 1967:85)Sotano de El Tigre, 24 km by road SW Jalpan, Mpo.de Landa de Matamoros (Reddell, 1979:100)San Joaquin RegionSotano de Herreras-see Grutas de San JoaquinCueva del Rincon, La Lagunita, 11 km SSW SanJoaquin, Mpo. de San Joaquin (San Joaquin,Fl4C58)Sotano del Rincon, La Lagunita, 11 km SSW SanJoaquin, Mpo. de San Joaquin (Mothes, 1978:63)(SanJoaquin,F14C58)Grutas de San Joaquin, 2 km SSW San Joaquin,2440 m, Mpo. de San Joaquin (Mothes, 1978:64-65) (San Joaquin, Fl4(58)San Juan RegionCave of the Bedding, 2 km N San Juan, Mpo. deArroyo Seco*Hoya de las Conchas, 3 km N San Juan, Mpo. deArroyo Seco (Stone and Jameson, 1977:28m, 2933, map)Sotano de Conchas-see Hoya de las ConchasCueva de Guayavitos, 1 km SW San Juan, Mpo. deArroyo Seco (Stone and Jameson, 1977:42)Cueva de la Milpa, 2 km W San Juan, Mpo. de ArroyoSeco*Sotano de Nogal, 5 km NW San Juan, Mpo. deArroyo Seco (Stone and Jameson, 1977:34m,35-39, map)Sotano de Pedregal, 1 km NE San Juan, Mpo. deArroyo Seco (Stone and Jameson, 1977:44m)El Sotanito, 1.5 km N San Juan, Ylpo. de ArroyoSeco (Stone and Jameson, 1977:20m, 21, map)Tancoyol RegionCueva del Agua del Rancho Ojo de Agua, 20 km NEJalpan, Mpo. de J alpanSotano de Lagunita Seca, Tancoyol, 20 km NE Jalpan,Mpo. de Jalpan*EI Socavon, Ojo de Agua, 20 km NE Jalpan, Mpo.de J alpan (Broussard, 1975:2m)Xilitla Plateau RegionCueva de Campamiento-see Sumidero del LlanoConejoCueva de los Grillos, 3 km N Cerro de la Luz, 13 kmW Xilitla, Mpo. de Llanda de MatamorosCueva de Llano de los Chiquitos-see Sumidero delLlano Chiquito*Sumidero del Llano Chiquito, 12 km W Xilitla, Mpo.de Landa de Matamoros (Fish, 1978b:41, 42m;Fish,1979:7)Cueva del Llano del Conejo-see Sumidero del LlanoConejo*Sumidero del Llano Conejo, 10 km ENE Xilitla,Mpo. de Landa de Matamoros (Fish, 1979:12,13m)302
Page 1:
A REVIEW OF THECAVERNICOLE FAUNA OF
Page 8:
. 1-----,, ,", ,\'"I/I~" )/Fig. I.-
Page 12 and 13:
collected a troglobitic cirolanid i
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C. Bolivar and J. Carranza investig
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springs in the vicinity of Cautro C
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Cueva del Diablo near Orizaba, Vera
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Those species of troglobite which h
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Table 2.-(continued)1970. Eodrilus
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CAVE REGIONSOnly the cave regions o
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Table 3.-(continued)TroglobitesTrog
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y the waters of the Amistad Reservo
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Although a few caves in the Norther
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an elevation of about 900 meters. T
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the gryllid cricket Paracophus suba
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has been Jiscussed by Segerstrom (1
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only apparent troglophile is the ps
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The cave fauna of this region is ri
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igneous rock immediately east of th
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Rio Soto la Marina and terminates a
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caves studied are many large stream
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unusual species is Neogovea mexasca
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pallidus Rowland, the nesticid spid
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troglobites: the trichoniscid isopo
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Cambridge, and Pholcophora bispinos
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of the genus Lutzomyia has not been
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The principal cavernous fonnation i
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ocks of Paleocene to Pliocene age.
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Beach and Supra-tidal Zone, the I\o
Page 66 and 67:
The aquatic fauna of Mexico and Cen
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oaxacae Hobbs from the Valle Nacion
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described, the genus occurs in cave
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large genus with numerous epigean s
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milliped, laniatorid harvestmen, an
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Table 4.-Summary of cave inhabiting
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Discussion.-This large, troglobitic
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any way a part of the true cave fau
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they have been washed in with soil
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part of the aquatic fauna. A very l
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The genus Helicodiscus is a signifi
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associated with true cave waters. P
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Family DaphnidaeWilson (1936) repor
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tul), Cueva de San Isidro, Cueva de
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30 m below entrance of Cueva de la
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underflow of rivers, alluvial grave
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the similarities between the distri
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Yucatan but did not identify the sp
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Protrichoniscus: Reddell, 1967a:24.
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It indudes five spccies, of which t
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Type-locality.-Pozzo, Etla, Oaxaca,
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Records.-Campeche: Volc
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164; Pearse, 1945:167, 168, fig. 13
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Family AtyidaeThe family Atyidae is
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Creaseria morleyi (Creaser)Blind sh
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the pond which is fed by the cave s
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philes. Pseudothelphusa (Pseudothel
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Typhlochactas elliotti MitchellTyph
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Distribution.-Known only from the t
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(Dumitresco and Orghidan, 1977); th
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Leucohya magnifica MuchmoreUndescri
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Discussion.-This is the largest spe
Page 131 and 132:
Type-Iocality.-S6tano de Huitzmolot
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Bartolo is a name applied to two ne
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Rowland, 1975b:19, 20; Dumitresco,
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Type-Iocality.-Actlin Kaua, Yucatan
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Distribution.-Known only from caves
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een collected. It is frequently fou
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Distribution.-Known from two caves
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actually a natural cave artificiall
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Discussion.-This large species has
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and T. patellaris Bryant from caves
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estricted to the Mexican Plateau (s
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maya Chamberlin and Ivie from Campe
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Metagonia chiquita GertschMetagonia
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Metagonia capilla: Mitchell and Kaw
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the cave and in general association
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DiscWlSion.-This is a delicate, eye
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Type-Iocality.-Cueva de Los Sabinos
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family. Clarence and Marie Goodnigh
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n. 2, San Luis Potosi; and K. rugos
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Type-locality.-Cueva de Apoala, San
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Records.-Chiapas: Sumidero de Canad
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Family HeterozerconidaeSpecimens of
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only one species. All known epigean
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nation must await study, but it is
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Mexico and Guatemala, and several u
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Type-locality.-Gruta de Atoyac, Ver
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Records.-San Luis PotOSI: Sotano de
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Records.-Veracruz: Grutas de Atoyac
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the genus Synoptura and removed the
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known from caves all contain troglo
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las Perlas, Cueva Chica de la Perra
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Discussion.-This species is distinc
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Speodesmus pecki ShearPolydesmidae,
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appears to be most closely related
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this is almost certainly M. russell
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Table 21.-Summary of cave inhabitin
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Type-Iocality.-Cueva Bonita del Pre
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Type-Iocality.-Cueva de El Jobo, El
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Order DipluraFamily CampodeidaeAlth
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of small soil inhabiting japygoids
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(Linnaeus) has been collected from
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Cave. Paracophus caecus appears to
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the most poorly known species in th
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Family EpipsocidaeThe family Epipso
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citata Van Tyne in Cenote de Piste,
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een found in caves and mines in Yuc
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ground dwelling habit, many beetles
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Type-Iocality.-Sotano de Tejamanil,
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Vigna Taglianti, 1972:118, 121-127,
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Veracruz) belongs in another subgen
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Troglobacanius sbordonii VomeroTrog
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Ptomaphagus (Adelops) is a group of
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(A.) speLaeus, from Grutas de Cacah
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epresented by many species. Tenebri
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Myodopsylla gentilis Jordan and Rot
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genus Pholeomyia have been collecte
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Wheeler (1938) has reported the pre
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263.264, 265, 267, 270, 273, fig. 5
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Anophthyctys jordani: Stefanelli, 1
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Prietella phreatophila Carranza, 19
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TyphlilJs: Pearse, 1938a:13, 15; Hu
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Family BufonidaeFour species of toa
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Oaxaca; and E. yucatanensis Lynch f
Page 258 and 259: sity of bats in the tropics. In add
Page 260 and 261: Three species of short-tailed bats
Page 262 and 263: abundant in the entrance area of Cu
Page 264 and 265: Andrews, E. W., IV. 1970. Balankanc
Page 266 and 267: Bolivar y Pieltain, C. 1940. Explor
Page 268 and 269: Brignoli, P. M. 1979a. The morpholo
Page 270 and 271: Conde, B. 1975. Description du prem
Page 272 and 273: Ediger, G. 1970. The Birthday Passa
Page 274 and 275: Gonzalez Ochoa, A. 1963b. Relacione
Page 276 and 277: Hobbs, H. H.,Jr., and T. C. Barr, J
Page 278 and 279: Jordan, c. B. 1937. Bringing in the
Page 280 and 281: McKenzie, D. 1965a. Caves of the Si
Page 282 and 283: Monod, T. 1975. Sur la distribution
Page 284 and 285: Peck, S. B., and J. J. Lewis. 1977.
Page 286 and 287: Reddell, J. R., and W. R. Elliott.
Page 288 and 289: Sanborn, C. C. 1936. Records and me
Page 290 and 291: Stone, B. 1979. The 1979 San Agusti
Page 292 and 293: Villa R., B. 1956. Una extrana y se
Page 294 and 295: Wimsatt, W. A. 1969. Transient beha
Page 296 and 297: +*Cueva de los Chivos, 5 km E Altam
Page 298 and 299: Sumidero Yochib, Yochib, 8 km NE Te
Page 300 and 301: Serrania del Burro RegionCave, Haci
Page 302 and 303: +Cueva de Tezoapa, 1.5 km E Acahuiz
Page 304 and 305: *Gruta Sur de San Bartolo, 9 km S S
Page 306 and 307: *Cueva del Puente de Fierro, 1.5 km
Page 310 and 311: Matehuala RegionSumidero 552, at Km
Page 312 and 313: *S6tano de Yerbaniz, 21.5 km NNE Ci
Page 314 and 315: TABASCO+Cave 2 km E Teapa, Mpo. de
Page 316 and 317: +Sinkhole, 12 m deep (No.2), Rancho
Page 318 and 319: Sotano de El Maguey, 5 km N Cuitlah
Page 320 and 321: *Cenote Aka Chen, 1 km E Tixcancal,
Page 322 and 323: *Cenote Xtacabiha, 9 km NNE Tikuch,
Page 324 and 325: ESCUINTLA+Cueva de los Ladrones, Fi
Page 326 and 327: Procambarus (Austrocambarus) oaxaca
Page 328 and 329: BelizeCayoMexobisilJm goodnighti-Ps
Page 330 and 331: Creaseria morleyi (Creaser) (p. 109
Page 332 and 333: Typhlias: De Buen, 1941 :9.Typhlias
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