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Metagonia capilla: Mitchell and Kawakatsu, 1973a:673 (error <strong>for</strong> M. pura); Reddell and Elliott,1973b:181 (error <strong>for</strong> M. pura).Type-Iocality.-Cueva de la Capilla, 13.5 km NWof Gomez Farias, Tamaulipas, Mexico.Distribution.-Known only from the type-locality.See Fig. 39.Discussion.-This pale, eyeless species is related toM. tlamaya.Metagonia tlamaya GertschMetagonia sp.: Reddell, 1967d:106; Reddell, 1971b:31 (Sotano de Tlamaya record only).Metagonia tlamaya Gertsch, 1971a:48, 83-84, 87,fig. liO-lil, 125; Reddell, 1973a:33, 39; Hoffmann,1976:25.Type.locality.~Sotano de Tlamaya, San Luis Potosi,Mexico.Distribution.-Known only from the type-locality.See Fig. 39.Discussion.-This pale, eyeless species is related toM. tinaja.Metagonia torete GertschMetagonia torete Gertsch, 1977b:105, 107,108-109,fig. 9, 25-26, 29-30; Reddell, 1977b:232, 240,241, 281, 292, 293.MEXICO4..4GUATEMALAii\i)/""_._._._.\r·,.....--,e/.//rFig. 41.-Distrihution of eight species of troglophilic pholcid spider of the genus Metagonia: 1, Metagonia coahuila; 2, M.suzanne; 3, M. pasquinii; 4, M. mcnatti; 5, M. maya; 6, M. yucatana; 7, Metagonia n. sp. 3; 8, M. blanda.152
Type-locality.-Cueva Sodzil, 5 km W Sucopo,Yucatan, Mexico.Distribution.-Known from caves in Campeche,Quintana Roo, and Yucatan. See Fig. 39.Records.-Yucatan: Cenote de Orizaba, Cueva deSodzil, Grutas de Tzab-Nah, and Cenote Xtacabiha.New records.-Campeche: Grutas de San Antonio;Quintana Roo: Cueva de Tancah (det. W. J. Gertsch).Discussion.-This is a small species with evanescenteyes and elongated legs.Pholcophora exigua GertschPholcophora exigua Gertsch, 1971a:80, 82, fig. 99100; Reddell, 1973a:39; Hoffmann, 1976:26;Reddell, 1977a:90.Type-locality.-Cueva de los Riscos, Sierra de laIndia, 6.5 km S of Mapimi, Durango, Mexico.Distribution.-Known only from the type-locality.See Fig. 42.Discussion.-This small, white species has the anteriormedian eyes obsolete and the lateral eyes reduced.It was collected from the remotest parts ofMEXICOGUATEMALAlb~//('\..... r".;.Fig. 42.-Distribution of troglobitic and troglophilic pholcid spiders of the genus Pholcophora: 1, Pholcophora exigua;2, P. mitchelli; 3, P. bolivari; 4, P. elliotti; 5, Pholcophora n. sp. 1; 6, Pholcophora n. sp. 2 and P. troglodyta; 7, Pholcophoran. spp. 3-4; 8, Pholcophora n. spp. 5-6; 9, Pholcophora n. sp. 7; 10, P. gruta; 11, P. bispinoso; 12, P. evansi; 13, Pholcophoran. sp. 8; 14, P. speophila; 15, P. pearsei; 16, P. quieta.153
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A REVIEW OF THECAVERNICOLE FAUNA OF
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. 1-----,, ,", ,\'"I/I~" )/Fig. I.-
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collected a troglobitic cirolanid i
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C. Bolivar and J. Carranza investig
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springs in the vicinity of Cautro C
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Cueva del Diablo near Orizaba, Vera
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Those species of troglobite which h
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Table 2.-(continued)1970. Eodrilus
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CAVE REGIONSOnly the cave regions o
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Table 3.-(continued)TroglobitesTrog
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y the waters of the Amistad Reservo
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Although a few caves in the Norther
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an elevation of about 900 meters. T
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the gryllid cricket Paracophus suba
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has been Jiscussed by Segerstrom (1
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only apparent troglophile is the ps
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The cave fauna of this region is ri
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igneous rock immediately east of th
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Rio Soto la Marina and terminates a
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caves studied are many large stream
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unusual species is Neogovea mexasca
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pallidus Rowland, the nesticid spid
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troglobites: the trichoniscid isopo
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Cambridge, and Pholcophora bispinos
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of the genus Lutzomyia has not been
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The principal cavernous fonnation i
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ocks of Paleocene to Pliocene age.
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Beach and Supra-tidal Zone, the I\o
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The aquatic fauna of Mexico and Cen
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oaxacae Hobbs from the Valle Nacion
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described, the genus occurs in cave
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large genus with numerous epigean s
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milliped, laniatorid harvestmen, an
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Table 4.-Summary of cave inhabiting
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Discussion.-This large, troglobitic
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any way a part of the true cave fau
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they have been washed in with soil
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part of the aquatic fauna. A very l
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The genus Helicodiscus is a signifi
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associated with true cave waters. P
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Family DaphnidaeWilson (1936) repor
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tul), Cueva de San Isidro, Cueva de
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30 m below entrance of Cueva de la
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underflow of rivers, alluvial grave
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the similarities between the distri
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Yucatan but did not identify the sp
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Protrichoniscus: Reddell, 1967a:24.
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It indudes five spccies, of which t
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Type-locality.-Pozzo, Etla, Oaxaca,
Page 108 and 109: Records.-Campeche: Volc
Page 110 and 111: 164; Pearse, 1945:167, 168, fig. 13
Page 112 and 113: Family AtyidaeThe family Atyidae is
Page 115 and 116: Creaseria morleyi (Creaser)Blind sh
Page 117 and 118: the pond which is fed by the cave s
Page 119 and 120: philes. Pseudothelphusa (Pseudothel
Page 121 and 122: Typhlochactas elliotti MitchellTyph
Page 123 and 124: Distribution.-Known only from the t
Page 125 and 126: (Dumitresco and Orghidan, 1977); th
Page 127 and 128: Leucohya magnifica MuchmoreUndescri
Page 129 and 130: Discussion.-This is the largest spe
Page 131 and 132: Type-Iocality.-S6tano de Huitzmolot
Page 133 and 134: Bartolo is a name applied to two ne
Page 135 and 136: Rowland, 1975b:19, 20; Dumitresco,
Page 137 and 138: Type-Iocality.-Actlin Kaua, Yucatan
Page 139 and 140: Distribution.-Known only from caves
Page 141 and 142: een collected. It is frequently fou
Page 143 and 144: Distribution.-Known from two caves
Page 146 and 147: actually a natural cave artificiall
Page 148 and 149: Discussion.-This large species has
Page 150 and 151: and T. patellaris Bryant from caves
Page 152 and 153: estricted to the Mexican Plateau (s
Page 154 and 155: maya Chamberlin and Ivie from Campe
Page 156 and 157: Metagonia chiquita GertschMetagonia
Page 160 and 161: the cave and in general association
Page 162 and 163: DiscWlSion.-This is a delicate, eye
Page 164 and 165: Type-Iocality.-Cueva de Los Sabinos
Page 166 and 167: family. Clarence and Marie Goodnigh
Page 168 and 169: n. 2, San Luis Potosi; and K. rugos
Page 170 and 171: Type-locality.-Cueva de Apoala, San
Page 172 and 173: Records.-Chiapas: Sumidero de Canad
Page 174 and 175: Family HeterozerconidaeSpecimens of
Page 176 and 177: only one species. All known epigean
Page 178 and 179: nation must await study, but it is
Page 180 and 181: Mexico and Guatemala, and several u
Page 182 and 183: Type-locality.-Gruta de Atoyac, Ver
Page 184 and 185: Records.-San Luis PotOSI: Sotano de
Page 186 and 187: Records.-Veracruz: Grutas de Atoyac
Page 188 and 189: the genus Synoptura and removed the
Page 190 and 191: known from caves all contain troglo
Page 192 and 193: las Perlas, Cueva Chica de la Perra
Page 194 and 195: Discussion.-This species is distinc
Page 196 and 197: Speodesmus pecki ShearPolydesmidae,
Page 198 and 199: appears to be most closely related
Page 200 and 201: this is almost certainly M. russell
Page 202 and 203: Table 21.-Summary of cave inhabitin
Page 204 and 205: Type-Iocality.-Cueva Bonita del Pre
Page 206 and 207: Type-Iocality.-Cueva de El Jobo, El
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Order DipluraFamily CampodeidaeAlth
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of small soil inhabiting japygoids
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(Linnaeus) has been collected from
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Cave. Paracophus caecus appears to
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the most poorly known species in th
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Family EpipsocidaeThe family Epipso
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citata Van Tyne in Cenote de Piste,
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een found in caves and mines in Yuc
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ground dwelling habit, many beetles
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Type-Iocality.-Sotano de Tejamanil,
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Vigna Taglianti, 1972:118, 121-127,
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Veracruz) belongs in another subgen
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Troglobacanius sbordonii VomeroTrog
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Ptomaphagus (Adelops) is a group of
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(A.) speLaeus, from Grutas de Cacah
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epresented by many species. Tenebri
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Myodopsylla gentilis Jordan and Rot
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genus Pholeomyia have been collecte
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Wheeler (1938) has reported the pre
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263.264, 265, 267, 270, 273, fig. 5
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Anophthyctys jordani: Stefanelli, 1
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Prietella phreatophila Carranza, 19
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TyphlilJs: Pearse, 1938a:13, 15; Hu
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Family BufonidaeFour species of toa
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Oaxaca; and E. yucatanensis Lynch f
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sity of bats in the tropics. In add
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Three species of short-tailed bats
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abundant in the entrance area of Cu
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Andrews, E. W., IV. 1970. Balankanc
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Bolivar y Pieltain, C. 1940. Explor
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Brignoli, P. M. 1979a. The morpholo
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Conde, B. 1975. Description du prem
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Ediger, G. 1970. The Birthday Passa
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Gonzalez Ochoa, A. 1963b. Relacione
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Hobbs, H. H.,Jr., and T. C. Barr, J
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Jordan, c. B. 1937. Bringing in the
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McKenzie, D. 1965a. Caves of the Si
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Monod, T. 1975. Sur la distribution
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Peck, S. B., and J. J. Lewis. 1977.
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Reddell, J. R., and W. R. Elliott.
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Sanborn, C. C. 1936. Records and me
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Stone, B. 1979. The 1979 San Agusti
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Villa R., B. 1956. Una extrana y se
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Wimsatt, W. A. 1969. Transient beha
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+*Cueva de los Chivos, 5 km E Altam
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Sumidero Yochib, Yochib, 8 km NE Te
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Serrania del Burro RegionCave, Haci
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+Cueva de Tezoapa, 1.5 km E Acahuiz
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*Gruta Sur de San Bartolo, 9 km S S
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*Cueva del Puente de Fierro, 1.5 km
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Cueva de los Riscos, 2 km S Rancho
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Cueva de El Coy-see Cueva del Nacun
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+*Cueva del Aire, 1.8 km at S85°W
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*Sistema Purificacion, Conrado Cast
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*Cueva del Nacimiento del Rio Frio,
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Sotano de Humo, 5 km NW San Andres,
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and Bohor, 1975:25-27, 26m; Reddell
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*Actun Xpukil, 3 km S Calcehtok, Mp
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Appendix 2LIST OF TROGLOBITES BY ST
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Paravachonium superbum-I'seudoscorp
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SUPPLEMENTWhile this manuscript was
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Cambala speobia (Chamberlin) (p. 19
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mida (Arachnida) in the New World.
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