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BelizeCayoMexobisilJm goodnighti-Pseudoscorpionida, HyidaeVachonilJm belizell.fe-Pseudoscorpionida, VachoniidaeMetagonia jarmila-Araneae, PholcidaeCynortina misteca-Opilionida, PhalangodidaeStygnomma pecki-Opilionida, PhalangodidaeJarmilka alba-Spirostreptida, Cambalidae322
SUPPLEMENTWhile this manuscript was in press several additionalpublications have appeared or been brought tomy attention which include infonnation on troglobitesfrom this region. The following citations followthe style used in the systematic review. The pagereferences in parentheses following the species namerefers to the page on which the species is mentioned.Bibliographic references follow the systematic citations.Diaptomus (Microdiaptomus) cokeri Osorio Tafall(p.81)Diaptomus cokeri: Bowman, 1979:225.Conilera stygia Packard (p. 84)Conilera stygia: Richardson, 1904:6.Conilera: Birstein, 1964:25.Creaseriella anops (Creaser) (p. 84)Creaseriella anops: Wilkens, 1979:265, 266, 267,fig. 2.Speocirolana bolivari (Rioja) (p. 86)Cirolana (part): Birstein, 1964:25.Speocirolana bolivari: Holt, 1973:223.Speocirolana peIaezi (Bolivar) (p. 86)Cirolana (part): Birstein, 1964:25.Speocirolana peIaezi: Holt, 1973:223.Caecidotea chiapas (Bowman) (p. 88)Caecidotea chiapas: Lewis, 1980:321,324.Caecidotea mitchelli Argano (p. 88)Caecidotea mitchelli: Lewis, 1980:324.Caecidotea pasquinii Argano (p. 88)Caecidotea pasquinii: Lewis, 1980:324.Caecidotea vomeroi Argano (p. 88)Caecidotea vomeroi: Lewis, 1980:324.Caecidotea zullinii Argano (p. 89)Caecidotea iullini: Lewis, 1980:324 (erroneousspelling).Mexistenasellus coahuila Cole and Minckley (p. 90)Mexistenasellus coahuila: Magniez, 1977:130.Mexistenasellus magniezi Argano (p. 90)Mexistenasellus sp. nov.: Magniez, 1977:130.Mexistenasellus parzefalli Magniez (p. 90)Mexistenasellus parzefalIi: Magniez, 1977:130.Mexistenasellus wilkensi Magniez (p. 90)Mexistenasellus wilkensi: Magniez, 1977:130.Brackenridgia bridgesi (Van Name) (p. 95)Protrichoniscus: Strouhal, 1953:175.Mayaweckelia cenoticoia Holsinger (p. 101)Mayaweckelia cenoticola: Wilkens, 1979:265, fig.2.Mayaweckelia spp. (part): Wilkens, 1979:266.Mayaweckelia yucatanensis Holsinger (p. 102)Mayaweckelia yucatanensis: Wilkens, 1979:265,266, fig. 2.Mayaweckelia spp. (part): Wilkens, 1979:266.Mexiweckelia colei Holsinger and Minckley (p. 102)Mexiweckelia (part): Holsinger and Longley, 1980:9, 10, 27, 51.Mexiweckelia mitchelli Holsinger (p. 102)Mexiweckelia (part): Holsinger and Longley, 1980:9, 10, 27, 51.Mexiweckelia particeps Holsinger (p. 102)Mexiweckelia particeps: Holsinger and Longley,1980:9, 10.Antromysis (Antromysis) cenotens&i Creaser (p. 103)Antromysis cenotensis: Wilkens, 1979:265, 266,fig. 2.Typhlatya campecheae Hobbs and Hobbs (p. 106)Typhlatya campecheae: Buden and Felder, 1977:114; Wilkens, 1979:265,266, fig. 2.Typhlatya (part): Wilkens, 1979:268,269.Typhlatya mitchelli Hobbs and Hobbs (p. 107)Typhlatya mitchelli: Buden and Felder, 1977:114; Wilkens, 1979:265,266,268, fig. 2.Typhlatya (part): Wilkens, 1979:268,269.Typhlatya pearsei (Creaser) (p. 107)Typhlatya pearsei: Buden and Felder, 1977:112; Wilkens, 1979:265,266,268,269, fig. 2.Typhlatya (part): Wilkens, 1979:268,269.Bithynops Iuscus Holthuis (p. 108)Bithynops: Strenth, 1978:67.323
Page 1:
A REVIEW OF THECAVERNICOLE FAUNA OF
Page 8:
. 1-----,, ,", ,\'"I/I~" )/Fig. I.-
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collected a troglobitic cirolanid i
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C. Bolivar and J. Carranza investig
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springs in the vicinity of Cautro C
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Cueva del Diablo near Orizaba, Vera
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Those species of troglobite which h
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Table 2.-(continued)1970. Eodrilus
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CAVE REGIONSOnly the cave regions o
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Table 3.-(continued)TroglobitesTrog
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y the waters of the Amistad Reservo
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Although a few caves in the Norther
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an elevation of about 900 meters. T
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the gryllid cricket Paracophus suba
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has been Jiscussed by Segerstrom (1
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only apparent troglophile is the ps
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The cave fauna of this region is ri
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igneous rock immediately east of th
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Rio Soto la Marina and terminates a
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caves studied are many large stream
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unusual species is Neogovea mexasca
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pallidus Rowland, the nesticid spid
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troglobites: the trichoniscid isopo
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Cambridge, and Pholcophora bispinos
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of the genus Lutzomyia has not been
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The principal cavernous fonnation i
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ocks of Paleocene to Pliocene age.
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Beach and Supra-tidal Zone, the I\o
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The aquatic fauna of Mexico and Cen
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oaxacae Hobbs from the Valle Nacion
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described, the genus occurs in cave
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large genus with numerous epigean s
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milliped, laniatorid harvestmen, an
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Table 4.-Summary of cave inhabiting
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Discussion.-This large, troglobitic
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any way a part of the true cave fau
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they have been washed in with soil
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part of the aquatic fauna. A very l
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The genus Helicodiscus is a signifi
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associated with true cave waters. P
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Family DaphnidaeWilson (1936) repor
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tul), Cueva de San Isidro, Cueva de
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30 m below entrance of Cueva de la
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underflow of rivers, alluvial grave
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the similarities between the distri
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Yucatan but did not identify the sp
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Protrichoniscus: Reddell, 1967a:24.
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It indudes five spccies, of which t
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Type-locality.-Pozzo, Etla, Oaxaca,
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Records.-Campeche: Volc
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164; Pearse, 1945:167, 168, fig. 13
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Family AtyidaeThe family Atyidae is
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Creaseria morleyi (Creaser)Blind sh
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the pond which is fed by the cave s
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philes. Pseudothelphusa (Pseudothel
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Typhlochactas elliotti MitchellTyph
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Distribution.-Known only from the t
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(Dumitresco and Orghidan, 1977); th
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Leucohya magnifica MuchmoreUndescri
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Discussion.-This is the largest spe
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Type-Iocality.-S6tano de Huitzmolot
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Bartolo is a name applied to two ne
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Rowland, 1975b:19, 20; Dumitresco,
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Type-Iocality.-Actlin Kaua, Yucatan
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Distribution.-Known only from caves
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een collected. It is frequently fou
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Distribution.-Known from two caves
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actually a natural cave artificiall
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Discussion.-This large species has
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and T. patellaris Bryant from caves
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estricted to the Mexican Plateau (s
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maya Chamberlin and Ivie from Campe
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Metagonia chiquita GertschMetagonia
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Metagonia capilla: Mitchell and Kaw
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the cave and in general association
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DiscWlSion.-This is a delicate, eye
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Type-Iocality.-Cueva de Los Sabinos
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family. Clarence and Marie Goodnigh
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n. 2, San Luis Potosi; and K. rugos
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Type-locality.-Cueva de Apoala, San
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Records.-Chiapas: Sumidero de Canad
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Family HeterozerconidaeSpecimens of
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only one species. All known epigean
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nation must await study, but it is
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Mexico and Guatemala, and several u
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Type-locality.-Gruta de Atoyac, Ver
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Records.-San Luis PotOSI: Sotano de
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Records.-Veracruz: Grutas de Atoyac
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the genus Synoptura and removed the
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known from caves all contain troglo
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las Perlas, Cueva Chica de la Perra
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Discussion.-This species is distinc
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Speodesmus pecki ShearPolydesmidae,
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appears to be most closely related
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this is almost certainly M. russell
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Table 21.-Summary of cave inhabitin
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Type-Iocality.-Cueva Bonita del Pre
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Type-Iocality.-Cueva de El Jobo, El
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Order DipluraFamily CampodeidaeAlth
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of small soil inhabiting japygoids
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(Linnaeus) has been collected from
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Cave. Paracophus caecus appears to
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the most poorly known species in th
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Family EpipsocidaeThe family Epipso
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citata Van Tyne in Cenote de Piste,
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een found in caves and mines in Yuc
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ground dwelling habit, many beetles
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Type-Iocality.-Sotano de Tejamanil,
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Vigna Taglianti, 1972:118, 121-127,
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Veracruz) belongs in another subgen
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Troglobacanius sbordonii VomeroTrog
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Ptomaphagus (Adelops) is a group of
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(A.) speLaeus, from Grutas de Cacah
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epresented by many species. Tenebri
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Myodopsylla gentilis Jordan and Rot
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genus Pholeomyia have been collecte
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Wheeler (1938) has reported the pre
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263.264, 265, 267, 270, 273, fig. 5
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Anophthyctys jordani: Stefanelli, 1
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Prietella phreatophila Carranza, 19
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TyphlilJs: Pearse, 1938a:13, 15; Hu
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Family BufonidaeFour species of toa
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Oaxaca; and E. yucatanensis Lynch f
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sity of bats in the tropics. In add
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Three species of short-tailed bats
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abundant in the entrance area of Cu
Page 264 and 265:
Andrews, E. W., IV. 1970. Balankanc
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Bolivar y Pieltain, C. 1940. Explor
Page 268 and 269:
Brignoli, P. M. 1979a. The morpholo
Page 270 and 271:
Conde, B. 1975. Description du prem
Page 272 and 273:
Ediger, G. 1970. The Birthday Passa
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Gonzalez Ochoa, A. 1963b. Relacione
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Hobbs, H. H.,Jr., and T. C. Barr, J
Page 278 and 279: Jordan, c. B. 1937. Bringing in the
Page 280 and 281: McKenzie, D. 1965a. Caves of the Si
Page 282 and 283: Monod, T. 1975. Sur la distribution
Page 284 and 285: Peck, S. B., and J. J. Lewis. 1977.
Page 286 and 287: Reddell, J. R., and W. R. Elliott.
Page 288 and 289: Sanborn, C. C. 1936. Records and me
Page 290 and 291: Stone, B. 1979. The 1979 San Agusti
Page 292 and 293: Villa R., B. 1956. Una extrana y se
Page 294 and 295: Wimsatt, W. A. 1969. Transient beha
Page 296 and 297: +*Cueva de los Chivos, 5 km E Altam
Page 298 and 299: Sumidero Yochib, Yochib, 8 km NE Te
Page 300 and 301: Serrania del Burro RegionCave, Haci
Page 302 and 303: +Cueva de Tezoapa, 1.5 km E Acahuiz
Page 304 and 305: *Gruta Sur de San Bartolo, 9 km S S
Page 306 and 307: *Cueva del Puente de Fierro, 1.5 km
Page 308: Cueva de los Riscos, 2 km S Rancho
Page 311 and 312: Cueva de El Coy-see Cueva del Nacun
Page 313 and 314: +*Cueva del Aire, 1.8 km at S85°W
Page 315 and 316: *Sistema Purificacion, Conrado Cast
Page 317 and 318: *Cueva del Nacimiento del Rio Frio,
Page 319 and 320: Sotano de Humo, 5 km NW San Andres,
Page 321 and 322: and Bohor, 1975:25-27, 26m; Reddell
Page 323 and 324: *Actun Xpukil, 3 km S Calcehtok, Mp
Page 325 and 326: Appendix 2LIST OF TROGLOBITES BY ST
Page 327: Paravachonium superbum-I'seudoscorp
Page 331 and 332: Cambala speobia (Chamberlin) (p. 19
Page 333: mida (Arachnida) in the New World.
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