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*S6tano de Yerbaniz, 21.5 km NNE Ciudad Valles,241.5 m, Mpo. de Ciudad Valles (Mitchell et al.,1977:55-56)*Hoya de Zimapan, 22 km NE Ciudad Valles, Mpo. deCiudad Valles (Morris, 1971:45; Russell, 1972c:139-140)Sierra de Alvarez RegionCave near San Francisco, Mpo. de Zaragoza (Elliottand Reddell, 1973:194) (Santa Catarina, F14A85)*Cave in Valle de los Fantasmas, 4 km NW San Francisco,Mpo. de Zaragoza (Elliott and Reddell,1973:194)(Santa Catarina, F14A85)S6tano in Valle de los Fantasmas, 4 km NW SanFrancisco, Mpo. de Zaragoza (Elliott and Reddell,1973:194) (Santa Catarina, F14A85)S6tano de Abernathy, Los S6tanos Unidos, 6 kmWNW San Francisco, Mpo. de Zaragoza (Elliott andReddell, 1973:192) (Santa Catarina, F14A85)S6tano de Araiia-see S6tano de las AraiiasSotano de las Araiias, Los S6tanos Unidos, 6 kmWNW San Francisco, Mpo. de Zaragoza (Elliott andReddell, 1973:192) (Santa Catarina, Fl4A85)Cueva de los Caballos, Los S6tanos Unidos, 6 kmWNW San Francisco, Mpo. de Zaragoza (Elliott andReddell, 1973:192 (Santa Catarina, F14A85)S6tano de Carlos, 2 km N San Francisco, Mpo. deZaragoza (Elliott and Reddell, 1973:192) (SantaCatarina, F14A85)Cueva de Carnicerias, NW San Francisco, Mpo. deZaragoza (Elliott and Reddell, 1973:192) (SantaCatarina, F14A85)Cueva de Cinquenta y Ocho, I km W Cinquenta yOcho and 4 km S San Francisco, Mpo. de Zaragoza(Elliott and Reddell, 1973:192) (Santa Catarina,Fl4A85)Cave of Dead Cow, 4 km S San Francisco, Mpo. deZaragoza (Santa Catarina, F14A85)Cueva de la Entrada Chica, Valle de los Fantasmas,4 km NW San Francisco, Mpo. de Zaragoza (Elliottand Reddell, 1973:192) (Santa Catarina, F14A85)Sumidero de Fantasmas, Valle de los Fantasmas,4 km NW San Francisco, Mpo. de Zaragoza (Elliottand Reddell, 1973:192) (Santa Catarina, F14A85)*SOtano de la Golondrina, Valle de los Fantasmas,4 km NW San Francisco, Mpo. de Zaragoza (Walsh,1972:50m, 70; Elliott and Reddell, 1973:192)(Santa Catarina, F14A85)S6tano de las Golondrinas-see S6tano de la GolondrinaCueva de la Iglesia, 1.6 km W San Francisco, Mpo. deZaragoza (Walsh, 1972:51m, 72; Elliott and Reddell,1973:192)(Santa Catarina, F14A85)Cueva de la Laguna, 6.5 km NNW San Francisco,Mpo. de Zaragoza (Elliott and Reddell, 1973:192193) (Santa Catarina, F14A85)Cueva de las Moscas, 1.5 km S San Francisco, Mpo.de Zaragoza (Elliott and Reddell, 1973:193)(Santa Catarina, F14A85)Cuevacita de Nopales, 1 km SW Cinquenta y Ochoand 5 km S San Francisco, Mpo. de Zaragoza(Santa Catarina, F14A85)Sotano de Nopales, 1 km SW Cinquenta y Ocho and5 km S San Francisco, Mpo. de Zaragoza (Walsh,1972:55m, 68; Elliott and Reddell, 1973:193)(Santa Catarina, F14A85)SOtano de Ojo de Agua, 9 km ESE San Francisco,Mpo. de Zaragoza (Walsh, 1972:54m, 67; Elliottand Reddell, 1973:193) (Santa Catarina, F14A85)Sotano de Pajaro, Los Sotanos Unidos, 6 km WNWSan Francisco, Mpo. de Zaragoza (Elliott andReddell, 1973:193)(Santa Catarina, Fl4A85)*Cueva de la Puente, 17 km SSE San Francisco, Mpo.de Zaragoza (Walsh, 1972:60-61, 63m; Elliott andReddell, 1973:193)(La Salitrera, Fl4C15)*S6tano de Puerto de los Lobos, 1.5 km S San Francisco,2340 m, Mpo. de Zaragoza (Walsh, 1972:56-57, 59m; Elliott and Reddell, 1973:193) (SantaCatarina, Fl4A85)Cueva de las Rusias, Las Rusias, 5 km E San Francisco,Mpo. de Zaragoza (Fish and Reddell, 1967:83) (Santa Catarina, Fl4A85)S6tano de San Francisco, San Francisco, 2200 m,Mpo. de Zaragoza (Elliott and Reddell, 1973:194;Anonymous, 1974c:4m) (Santa Catarina, F14A85)*S6tano de San Francisco n. 2, 0.5 km S San Francisco,2300 m, Mpo. de Zaragoza (Elliott andReddell, 1973:194) (Santa Catarina, Fl4A85)S6tano de Super-Macho, Los Sotanos Unidos, 6 kmWNW San Francisco, Mpo. de Zaragoza (Elliottand Reddell, 1973:194) (Santa Catarina, Fl4A85)Sierra EI Pino RegionCueva de Los Avalos, Ejido de Los Avalos, 14 kmENE Ciudad del Maiz, 1140 m, Mpo. de Ciudad delMaiz (Reddell, 1965d:49) (Ciudad del Maiz,Fl4A78)Cueva Seca de Los Avalos, Ejido de Los Avalos,14 km ENE Ciudad del Maiz, 1180 m, Mpo. deCiudad del Mafz (Reddell, 1965d:49) (Ciudad delMaiz, F14A78)*Cueva de La Lagunita, La Lagunita, 16 km NE Ciudaddel Maiz, 1150 m, Mpo. de Ciudad del Maiz(Reddell, 1965d:49; Reddell, 1973g:96) (Ciudaddel Maiz, Fl4A78)Cueva de Mosca, La Lagunita, 16 km NE Ciudad delMafz, 1150 m, Mpo. de Ciudad del Mafz (Reddell,1979:101) (Ciudad del Maiz, F14A78)Cueva de los Murcielagos, 15 km NE Ciudad delMaiz, Mpo. de Ciudad del Maiz (Ciudad del Maiz,F14A78)Cueva de los Muros, 15 km NE Ciudad del Maiz, Mpo.de Ciudad del Maiz (Ciudad del Mafz, Fl4A78)Xilitla RegionS6tano near Tlamaya, 2.5 km NNW Xilitla, Mpo. deXilitla+Cave 4 km W on road to Xilitla, Mpo. de Xilitla(Pine, 1972:41)+Cave near road between Pan American highway andXilitla, Mpo. de Xilitla (Zweifel, 1956:16-17)(NOTE: This is probably Cueva de El Jobo)+Large cave 9 km NNE Xilitla, Mpo. de Xilitla (Dalquest,195'3a:25, 30)+Cueva del Ahuate n. 1, Puerto del Ahuate, 2 km SWXilitla, 721 m, Mpo. de Xilitla (Bonet, 1953a:57,pl.5m)*Cueva del Ahuate n. 2, Puerto del Ahuate, 2 km SWXilitla, 721 m, Mpo. de Xilitla (Bonet, 1953a:5868, pI. 5m)306
+*Cueva del Aire, 1.8 km at S85°W from church ofHuichihuayan, 190 m, Mpo. de Xilitla (Bonet,1953a:38-42,39m)+*Cueva de la Barranca, near La Barranca, 1 km NEAhuacatIan, 969 m, Mpo. de Xilitla (Bonet, 1953a:72-76, pi. 7m)Cuevita del Camino, Xilitlilla, 5 km E Ahuacatlan,Mpo. de XilitlaCueva de Campamiento-see Sumidero del LlanoConejo, QUERETAROCueva de la Cisterna, on trail to La Silleta fromTlamaya, NW Xilitla, 1330 m, Mpo. de XilitlaCueva de Cristian, 4 km E Xilitla, Mpo. de Xilitla+*Cueva de los Cuchos, near Plan de Juarez, 2 km SXilitla, 615 m, Mpo. de Xilitla (Bonet, 1953a:6872,69m)Cueva de la Gorra-see Cueva de la PorraCueva de los Grillos-see QUERETARO+*Cueva de la Hoya, 1 km NE Ahuacatlan, 938 m,Mpo. de Xilitla (Bonet, 1953a:83-86, pi. 9m)Sotano de las Hoyas, 1 km NW Ahuacatlan, Mpo. deXilitla (Walsh, 1972:109)Cueva de Huichihuayan-see Cueva del Nacimientodel Rio Huichihuayan*Sotano de Huitzmolotitla, 1 km ESE Tlamaya, 2 kmNNW Xilitla, Mpo_ de Xilitla (Russell and Raines,1967:90)Cueva de la Iglesia, AhuacatIan, Mpo. de Xilitla (Fishand Reddell, 1967:84)*Cueva de EI Jobo, 5 km E XilitIa, 445 m, Mpo. deXilitIa (Bonet, 1953a:44-50, 46m)*Cueva de la Laja, 0.75 km NE AhuacatIan, 870 m,Mpo. de Xilitla (Bonet, 1953a:76-82, 17m)Cueva de Llano de los Chiquitos-see Sumidero delLlano Chiquito, QUERETAROSumidero del Llano Chiquito-see QUERETAROCueva del Llano del Conejo-see Sumidero del LlanoConejo, QUERETAROSumidero del Llano Conejo-see QUERETAROCueva del Lobo-see Cueva de EIJobo+Cueva de la Mujer del Agua, 1.8 km at S85° W fromchurch of Huichihuayan, 245 m, Mpo. de XilitIa(Bonet, 1953a:42-44, 43m)Cueva del Nacimiento del Rio Huichihuayan, 2 kmSW Huichihuayan, lIO m, Mpo. de Xilitla (Bonet,1953a:36-38)*S6tano de la Navidad, SW of La Silleta, NW Xilitla,1970 m, Mpo. de XilitlaCueva de la Parra-see Cueva de la PorraCueva de Poca Ventana-see Cueva del Ahuate n. 2*Cueva de la Porra, 1 km SW Tlamaya, 2 km NNWXilitIa, Mpo. de Xilitla (Russell and Raines, 1967:94)Sotano de la Porra, 1 km SW Tlamaya, 2 km NNWXilitla, Mpo. de Xilitla (Bridgemon, 1974:111,143m)*Cueva de Potrerillos, Rancho de Potrerillos, 2 kmWSW Ahuacatlan, 1250 m, Mpo. de Xilitla (Bonet,1953a:86-91, pi. 10m)SOtano de Potrerillos, Rancho de Potrerillos, 2 km WAhuacatlan, Mpo. de Xilitla (Reddell, 1979:100)*SOtano de Potrero, 1 km N Ahuacatlan, Mpo. deXilitla*SOtano del Pozo, 1 km W AhuacatIan, Mpo. de Xilitla(Russell and Raines, 1967:99, 102m)Sotano del Rancho de la Barranca, 5 km NNE Ahuacatlan,Mpo. de Xilitla (Reddell, 1973d:87)*Cueva del Salitre, 0.5 km NNE Xilitla, 460 m, Mpo.de Xilitla (Bonet, 1953a:50, 52-55, 51m; Russelland Raines, 1967:95, 98m)*Cueva de Ia Selva, 2 km SW Xilitla, Mpo. de Xilitla(Russell and Raines, 1967:99, 101m)*Sotano de Tlamaya, Tlamaya, 2.5 km NNW Xilitla,Mpo. de Xilitla (Russell and Raines, 1967:90, 92m,94)Cueva del Xobo-see Cueva de EI JoboUnplaced CavesSINALOA+Cave near San Luis Potosi (Gonzalez-Ochoa, 1963a:67)+Cueva del Platanito (Villa R., 1967:5)+Large cavern system 1.5 km SW Panuco, Mpo. de Concordia(Bateman and Vaughan, 1974:45,52)+Small cave near La Cueva Chinacatera, 23 km W Pericos,Mpo. de Mocorito (Constantine, 1967:18)+Shallow caves 16 km NNW Los Mochis, Mpo. de LosMochis (Jones et a1., 1972:5)+Cave, Santa Lucia, Mpo. de Concordia (Jones et a1.,1972:7)+Cave 2.5 km NW Topolobampo, Mpo. de Los Mochis(.Jones et aI., 1972:5)+Cave 1.5 km N Zaragoza, Mpo. de Los Mochis (Jones etaI., 1972:13)+Cueva de la Chinacatera, 23 km W Pericos, Mpo. dc Mocorito(Constantine, 1967:18)+Cueva de Don Cristino, 20 km E Mazatlan, Mpo. deMazatlan (Villa R., 1967:152)Cueva de Monte Largo-see Cueva de la ChinacateraSONORA+Large cave, Chinobampo, Mpo. dc Navojoa (Burt, 1938:19)+Shallow caves, Chinobampo, Mpo. de Navojoa (Burt,1938:20)+Cave 24 km NW Guaymas, Mpo. de Guaymas (Burt,1938:19)+Cave, Guiricoba, Mpo. de Alamos (Burt, 1938:21)+Cave 1.5 km W Mayterrena (=Mayorena), 15 km Empalme,Mpo. de Empalme (Bradshaw and Hayward,1960:282)+Cave, PHares de Nacozari, Mpo. de Nacozari (Burt,1938:21)+Cave near San Bernardo, Mpo. de Alamos (Burt andHooper, 1941:2)+Cave below Santa Maria Mine, near EI Tigre, Mpo. de?Naocozari (Burt, 1938:21)+Mina La Aduana, 8 km W Alamos, Mpo. de Alamos(Horst, 1972:49-52, 51m)Cueva de Carbo-see Cueva del TigreLas Minas Cucaracha-see Mina La Aduana+Cueva de la Higuera, San Miguel de Horcasitas, Mpo.de San Miguel de Horcasitas (Roth, 1968:28)+Cueva del Tigre, 22 km SSE Carbo, Mpo. de Carbo(Mitchell,1965:568-577)307
Page 1:
A REVIEW OF THECAVERNICOLE FAUNA OF
Page 8:
. 1-----,, ,", ,\'"I/I~" )/Fig. I.-
Page 12 and 13:
collected a troglobitic cirolanid i
Page 14 and 15:
C. Bolivar and J. Carranza investig
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springs in the vicinity of Cautro C
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Cueva del Diablo near Orizaba, Vera
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Those species of troglobite which h
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Table 2.-(continued)1970. Eodrilus
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CAVE REGIONSOnly the cave regions o
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Table 3.-(continued)TroglobitesTrog
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y the waters of the Amistad Reservo
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Although a few caves in the Norther
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an elevation of about 900 meters. T
Page 35 and 36:
the gryllid cricket Paracophus suba
Page 37 and 38:
has been Jiscussed by Segerstrom (1
Page 39 and 40:
only apparent troglophile is the ps
Page 41 and 42:
The cave fauna of this region is ri
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igneous rock immediately east of th
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Rio Soto la Marina and terminates a
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caves studied are many large stream
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unusual species is Neogovea mexasca
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pallidus Rowland, the nesticid spid
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troglobites: the trichoniscid isopo
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Cambridge, and Pholcophora bispinos
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of the genus Lutzomyia has not been
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The principal cavernous fonnation i
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ocks of Paleocene to Pliocene age.
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Beach and Supra-tidal Zone, the I\o
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The aquatic fauna of Mexico and Cen
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oaxacae Hobbs from the Valle Nacion
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described, the genus occurs in cave
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large genus with numerous epigean s
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milliped, laniatorid harvestmen, an
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Table 4.-Summary of cave inhabiting
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Discussion.-This large, troglobitic
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any way a part of the true cave fau
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they have been washed in with soil
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part of the aquatic fauna. A very l
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The genus Helicodiscus is a signifi
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associated with true cave waters. P
Page 90 and 91:
Family DaphnidaeWilson (1936) repor
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tul), Cueva de San Isidro, Cueva de
Page 94 and 95:
30 m below entrance of Cueva de la
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underflow of rivers, alluvial grave
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the similarities between the distri
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Yucatan but did not identify the sp
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Protrichoniscus: Reddell, 1967a:24.
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It indudes five spccies, of which t
Page 106 and 107:
Type-locality.-Pozzo, Etla, Oaxaca,
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Records.-Campeche: Volc
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164; Pearse, 1945:167, 168, fig. 13
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Family AtyidaeThe family Atyidae is
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Creaseria morleyi (Creaser)Blind sh
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the pond which is fed by the cave s
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philes. Pseudothelphusa (Pseudothel
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Typhlochactas elliotti MitchellTyph
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Distribution.-Known only from the t
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(Dumitresco and Orghidan, 1977); th
Page 127 and 128:
Leucohya magnifica MuchmoreUndescri
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Discussion.-This is the largest spe
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Type-Iocality.-S6tano de Huitzmolot
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Bartolo is a name applied to two ne
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Rowland, 1975b:19, 20; Dumitresco,
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Type-Iocality.-Actlin Kaua, Yucatan
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Distribution.-Known only from caves
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een collected. It is frequently fou
Page 143 and 144:
Distribution.-Known from two caves
Page 146 and 147:
actually a natural cave artificiall
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Discussion.-This large species has
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and T. patellaris Bryant from caves
Page 152 and 153:
estricted to the Mexican Plateau (s
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maya Chamberlin and Ivie from Campe
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Metagonia chiquita GertschMetagonia
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Metagonia capilla: Mitchell and Kaw
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the cave and in general association
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DiscWlSion.-This is a delicate, eye
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Type-Iocality.-Cueva de Los Sabinos
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family. Clarence and Marie Goodnigh
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n. 2, San Luis Potosi; and K. rugos
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Type-locality.-Cueva de Apoala, San
Page 172 and 173:
Records.-Chiapas: Sumidero de Canad
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Family HeterozerconidaeSpecimens of
Page 176 and 177:
only one species. All known epigean
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nation must await study, but it is
Page 180 and 181:
Mexico and Guatemala, and several u
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Type-locality.-Gruta de Atoyac, Ver
Page 184 and 185:
Records.-San Luis PotOSI: Sotano de
Page 186 and 187:
Records.-Veracruz: Grutas de Atoyac
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the genus Synoptura and removed the
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known from caves all contain troglo
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las Perlas, Cueva Chica de la Perra
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Discussion.-This species is distinc
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Speodesmus pecki ShearPolydesmidae,
Page 198 and 199:
appears to be most closely related
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this is almost certainly M. russell
Page 202 and 203:
Table 21.-Summary of cave inhabitin
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Type-Iocality.-Cueva Bonita del Pre
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Type-Iocality.-Cueva de El Jobo, El
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Order DipluraFamily CampodeidaeAlth
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of small soil inhabiting japygoids
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(Linnaeus) has been collected from
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Cave. Paracophus caecus appears to
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the most poorly known species in th
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Family EpipsocidaeThe family Epipso
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citata Van Tyne in Cenote de Piste,
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een found in caves and mines in Yuc
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ground dwelling habit, many beetles
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Type-Iocality.-Sotano de Tejamanil,
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Vigna Taglianti, 1972:118, 121-127,
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Veracruz) belongs in another subgen
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Troglobacanius sbordonii VomeroTrog
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Ptomaphagus (Adelops) is a group of
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(A.) speLaeus, from Grutas de Cacah
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epresented by many species. Tenebri
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Myodopsylla gentilis Jordan and Rot
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genus Pholeomyia have been collecte
Page 244 and 245:
Wheeler (1938) has reported the pre
Page 246 and 247:
263.264, 265, 267, 270, 273, fig. 5
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Anophthyctys jordani: Stefanelli, 1
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Prietella phreatophila Carranza, 19
Page 252 and 253:
TyphlilJs: Pearse, 1938a:13, 15; Hu
Page 254 and 255:
Family BufonidaeFour species of toa
Page 256 and 257:
Oaxaca; and E. yucatanensis Lynch f
Page 258 and 259:
sity of bats in the tropics. In add
Page 260 and 261:
Three species of short-tailed bats
Page 262 and 263: abundant in the entrance area of Cu
Page 264 and 265: Andrews, E. W., IV. 1970. Balankanc
Page 266 and 267: Bolivar y Pieltain, C. 1940. Explor
Page 268 and 269: Brignoli, P. M. 1979a. The morpholo
Page 270 and 271: Conde, B. 1975. Description du prem
Page 272 and 273: Ediger, G. 1970. The Birthday Passa
Page 274 and 275: Gonzalez Ochoa, A. 1963b. Relacione
Page 276 and 277: Hobbs, H. H.,Jr., and T. C. Barr, J
Page 278 and 279: Jordan, c. B. 1937. Bringing in the
Page 280 and 281: McKenzie, D. 1965a. Caves of the Si
Page 282 and 283: Monod, T. 1975. Sur la distribution
Page 284 and 285: Peck, S. B., and J. J. Lewis. 1977.
Page 286 and 287: Reddell, J. R., and W. R. Elliott.
Page 288 and 289: Sanborn, C. C. 1936. Records and me
Page 290 and 291: Stone, B. 1979. The 1979 San Agusti
Page 292 and 293: Villa R., B. 1956. Una extrana y se
Page 294 and 295: Wimsatt, W. A. 1969. Transient beha
Page 296 and 297: +*Cueva de los Chivos, 5 km E Altam
Page 298 and 299: Sumidero Yochib, Yochib, 8 km NE Te
Page 300 and 301: Serrania del Burro RegionCave, Haci
Page 302 and 303: +Cueva de Tezoapa, 1.5 km E Acahuiz
Page 304 and 305: *Gruta Sur de San Bartolo, 9 km S S
Page 306 and 307: *Cueva del Puente de Fierro, 1.5 km
Page 308: Cueva de los Riscos, 2 km S Rancho
Page 311: Cueva de El Coy-see Cueva del Nacun
Page 315 and 316: *Sistema Purificacion, Conrado Cast
Page 317 and 318: *Cueva del Nacimiento del Rio Frio,
Page 319 and 320: Sotano de Humo, 5 km NW San Andres,
Page 321 and 322: and Bohor, 1975:25-27, 26m; Reddell
Page 323 and 324: *Actun Xpukil, 3 km S Calcehtok, Mp
Page 325 and 326: Appendix 2LIST OF TROGLOBITES BY ST
Page 327 and 328: Paravachonium superbum-I'seudoscorp
Page 329 and 330: SUPPLEMENTWhile this manuscript was
Page 331 and 332: Cambala speobia (Chamberlin) (p. 19
Page 333: mida (Arachnida) in the New World.
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