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searchable PDF - Association for Mexican Cave Studies

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Speodesmus pecki ShearPolydesmidae, undescribed genus and species: Reddelland Mitchell, 1971a:150.Speodesmus pecki Shear, 1974:241, 272, 273-274,fig. 55-58; Shear, 1977a:246, 247.Type-Iocality.-Cueva del Pachon, 12 mi south ofCiudad Mante, Tamaulipas, Mexico.Distribution.-Known only from the type-locality.See Fig. 54.Discussion.-Shear (1974) placed this species inSpeodesmus with the admission that it may be representativeof an undescribed genus. Speodesmus isknown only by troglobitic species in Central Texasand southern New Mexico. An undescribed speciesoccurs in Grutas del Palmito, Nuevo Leon.Tylogoneus minus CauseyTylogoneus minus Causey, 1973:121, fig. 32; Shear,1977a:246.Type-Iocality.-Cueva de Tres Manantiales (=Cuevadel Ojo de Agua de Manantiales), Sierra de Guatemala,17 km SW Gomez Farias, Tamaulipas, Mexico.Distribution.-Known only from the type-locality.See Fig. 54.Discussion.-This species is known only by a singlemale and two fragmentary specimens of each sex. Thegenus Tylogoneus is known only by the two troglobiteslisted here. Shear (1977a) reported a female ofthis genus from Sumidero de Canada, Chiapas.Tylogoneus rainesi CauseyTylogoneus rainesi Causey, 1973:121, fig. 27-31;Shear, 1977a:246, 247.Type-Iocality.-Cueva de Poca Ventana (=Cuevadel Ahuate n. 2), Xilitla, San Luis Potosi, Mexico.Distribution.-Known only from the type-locality.See Fig. 54.Discussion.-This species is known only from themale holotype.Family TridontomidaeTridontomus loomisi ShearTridontomus loomisi Shear, 1977a:241-243, fig. 14­18.Type-locality.-Cueva Seamay, near Senahu, AltaVerapaz, Guatemala.Distribution.-Known only from the type-locality.See Fig. 54.Discussion.-This species is known only from themale holotype. The family Tridontomidae includesonly two species in addition to this troglobite. Theother two species, Tridontomus procerus Loomis andHoffman and Aenigmopus alatus Loomis and Hoffmanare known from Sepacuite and Seamay, AltaVerapaz, Guatemala, respectively. All three specieshave very long legs and antennae and weakly sclerotizedcuticles. Shear (1977a) speculates that the twospecies described by Loomis and Hoffman (1962)are also from caves since well known caves occur atboth Sepacuite (Cueva Sepacuite) and Seamay (CuevaSeamay).Family XystodesmidaeThe family Xystodesmidae is among the more conspicuouselements in the <strong>for</strong>est fauna of Mexico. Thisfamily has, however, not been successful in the colonizationof caves, and none are known to be troglobitesor troglophiles. The genera Cruzodesmus andRhysodesmus have both been collected from manycaves in the <strong>for</strong>ested, mountain regions of Mexico.These millipeds all appear either to have entered cavesaccidentally with washed-in debris, to have fallen intovertical cave entrances, or to have utilized the moistcave entrances <strong>for</strong> temporary shelter.Order PolyzoniidaFamily PolyzoniidaeThis family is represented in <strong>Mexican</strong> caves onlyby an undetermined species of the genus Siphonotusin Cueva de la Mina, Tamaulipas. It is presumably anaccidental.Family SiphonophoridaeOne species of siphonophorid, Siphonophora sabachanaChamberlin, has been described from <strong>Mexican</strong>caves. It was found in Actun Sabaca, Yucatan, and islisted by Nicholas (1962) as a troglobite, but isclearly an accidental or trogloxene. It has recentlybeen collected in Cueva Luchil, Yucatan. This or arelated species has also been taken from three otherYucatlin caves.Order SpirobolidaFamily AtopetholidaeThe only species of atopetholid identified fromcaves in Mexico is Aztecolus nigrior (Chamberlin).This species was found below the entrance drop intoChevy Sink, Nuevo Leon.Family MessicobolidaeThis family is represented only by undeterminedspecies of the genera Anelus and Messicobolus from190

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