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searchable PDF - Association for Mexican Cave Studies

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Discussion.-This eyeless species is apparently restrictedin its distribution to caves near Xilitla. Bothof the caves from which it is recorded are deep wetsystems.Glomeroides pellucidus ShearGlomeroides pellucidus Shear, 1974:248-249, fig.5-8.Type-Iocality.-Cueva del Ojo de Agua Grande,Paraje Nueva (=Nuevo), Cordoba, Veracruz, Mexico.Distribution.-Known only from the type-locality.See Fig. 52.Discussion.-This species possesses cuticular remnantsof ocelli. A single specimen was collected recentlyfrom silt in an upper level.Glomeroides promiscus CauseyGlomeroides promiscus Causey, 1964b:63, 65-66,fig. 6-8; Reddell, 1966a:13; Reddell, 1967c:55;Loomis, 1968:7; Reddell, 1971b;43; Reddell andMitchell, 1971b:189; Reddell, 1973a:34, 40;Reddell and Elliott, 1973b:181, 185; Shear, 1974;248,249; Vomero, 1974:350.Type-Iocality.-Sotano de Gomez Farias, 5 kmeast of Gomez Farias, Tamaulipas, Mexico.Distribution.-Known only from caves in theSierra de Guatemala, Tamaulipas. See Fig. 52.Records.-Tamaulipas: Sotano de Gomez Farias,?Cueva de la Mina, Sotano del Naranjo, Cueva delOjo de Agua de Manantiales, ?Cueva del· Rancho delCielo n. 3, and ?Cueva del Rancho del Cielo n. 7.Discussion.-This species possesses a few ocelli butis doubtless a troglobite. The records indicated by aquestion mark are represented only by females andare thus tentative. Immature specimens taken inCueva del Nacimiento del Rio Frio, Tamaulipas,probably also belong to this species. This milliped isusually found on moist silt and frequently in directassociation with other species of milliped.Order ChordeumidaThe order Chordeumida is the dominant group inthe milliped fauna of the caves of the eastern UnitedStates. It has not been nearly so successful in thecaves of Mexico and Central America; few specieshave invaded caves and fewer yet have become troglobites.Only two families in this order are representedin the cave fauna of Mexico.Family CleidogonidaeShear (1972) has monographed the family Cleidogonidaeand reviewed the order Chordeumida inNorth America. He reported six species of Cleidogonafrom caves in Mexico; two additional species havebeen added since (Shear, 1974, 1977a). None of thesespecies shows the high degree of adaptation to thecave environment which is present in some othermilliped families in Mexico, but five appear to beadapted to cave life and are included here (see Fig.53). Of the remaining three, one (c. crystallina Shear)is known only from caves and may now be isolated inthe cave environment. The two other species are alsoknown from epigean localities. Cleidogona mayapecShear has been identified from Cueva del Judio, Queretaro,and C. totonaca Shear has been found in SOtanodel Gobernador, Queretaro. Other collections ofthis family from caves in Tamaulipas and Veracruzawait study.Cleidogona baroqua ShearCleidogona baroqua Shear, 1972:198,207,209-210,fig. 197-200.Type-Iocality.-Sotano de San Augustin (=Agustin),Huautla de Jimenez, Oaxaca, Mexico.Distribution.-Known only from the type-locality.See Fig. 53.Discussion.-This depigmented species possessesocelli and is only tentatively considered to be a troglobite.It is a member of the crucis group. This speciesgroup includes the troglobitic C. crucis (Chamberlin)from Veracruz and three epigean species fromOaxaca. Shear (1972) considers this to be the mostprimitive group in the genus. Because of the presenceof ocelli, this species would have to be considered avery recent troglobite. The epigean species of thecrucis group are known from comparatively highelevations (1700 to 2500 meters), whereas C. baroquais found at about 1400 meters. The more highlyevolved troglobite C. crucis is found at 500 meters.Although only much additional collecting on the surfaceand in caves can clarify the situation, it is possiblethat the epigean ancestor of C. crucis becameextinct in the Veracruz lowlands much earlier thanin the Huautla area and that it or its closest relativesstill exist at high elevations in Oaxaca. This may be asimilar situation to that in the scorpion genus Typhlochactasand the milliped genus Mexiterpes.Cleidogona crucis (Chamberlin)Cavota crucis Chamberlin, 1942:8-9, fig. 3-7; Chamberlin,1943b;5, 36; Bonet, 1946a:112; Nicholas,1962:177; Loomis, 1968:66; Reddell, 1971a:222;Reddell, 1971b;44.Cleidogona crucis: Shear, 1972:207, 209, fig. 193­196; Shear, 1974:245,259.175

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