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TABASCO+Cave 2 km E Teapa, Mpo. de Teapa (Winkelmann,1962:112)+Cave 3.5 km by road NNE Teapa, Mpo. de Teapa (VillaR.,1967:210)+Cave 4.4 km SE Teapa, Mpo. de Teapa (Lay, 1962:374,375)+Well 22 km SE Villahennosa, Mpo. de Villahennosa(Zullini,1977:76)*Cueva del Azufre, 3.5 km S Tapijulapa, 50 m, Mpo. deTatotalpa (Gordon and Rosen, 1962:360-362, 361m;Sbordoni et al., 1974:15-16)Cueva de El Balneario, 3.5 km NE Teapa, 40 m, Mpo. deTeapa*Grutas del Cocomi, 3 km NE Teapa, 40 m, Mpo. deTeapa (Sbordoni et al., 1974:14-15)*Resumidero del Cocona, 3 km NE Teapa, 40 m, Mpo. deTeapa+Cueva de Don Luis, Finca Rosarita, 3.3 km NE, 3 km E,or SE Teapa, Mpo. de Teapa (Wimsatt, 1969:234)+Cueva del Escorpion, 5 km NNE Teapa, Mpo. de Teapa(Villa R., 1967:149)+Cueva de la Frontera, 1 km ESE El Azufre, on boundarybetween Tabasco and Chiapas, 40 m, Mpo. de Teapa(Sbordoni et al., 1977:21, 22m)+"La Gruta," Ejido Ignacio Allende, 20 0NE El Azufre onboundary between Tabasco and Chiapas, Mpo. deTeapa (Villa R., 1967:157.178)+Cueva La Murcielaguera, 5.3 km by road NNE Teapa,Mpo. de Teapa (Handley, 1966:300)+Cueva del Rancho El Azufre, 8 km W Teapa, Mpo. deTeapa (Villa R., 1967:327)TAMAULIPASAltas Cumbres RegionCueva Aire Mal, 15 km SSW Ciudad Victoria, 1350 m,Mpo. de Ciudad Vict9ria (Chabert, 1973:21, 23m,24) (Ciudad Victoria, F14A29)Sotano de Altas Cumbres n. I-see Cueva BonitaSotano de Altas Cumbres n. 2-see Cueva Aire MalCueva Bonita, 15 km SSW Ciudad Victoria, 1350 m,Mpo. de Ciudad Victoria (Chabert, 1973:21, 22m)(Ciudad Victoria, F14A29)+Cueva La Mula, 10 km W Joya Verde, 28 km SWCiudad Victoria, 800 m, Mpo. de Ciudad Victoria(Alvarez, 1963:339) (Ciudad Victoria, Fl4A29)Cerro El Aire RegionSotano de Abasolo, Abasolo, Mpo. de AbasoloCerro Gordo Region+Cave 3 km WNW EI Carrizo, 14 km SW Llera deCanales, Mpo. de Llera de Canales (Alvarez, 1963:405)(Gomez Farias, FI4A49)La Libertad RegionCueva de "El Murcielago"-see Cueva de Los TronconesCueva de la Presa-see Cueva de Los Troncones+Cueva de la Sepultura, 6 km NW Ciudad Victoria,740 m, Mpo. de Ciudad Victoria (Malaga Alba andVilla R., 1957:539, 543, 545, 567 (Guemez,Fl4A19)+Cueva de Los Troncones, near La Libertad, 8 km NWCiudad Victoria, Mpo. de Ciudad Victoria (RodriguezCabo, 1953:361-362) (Guemez, Fl4A19)Purificacion Region+Cave, El Chihue, 31 km NW Ciudad Victoria, 1800 m,Mpo. de Villa Hidalgo (Walker, 1955a:2) (CasasReales, Fl4A18)*Small spring 0.5 km W Comado Castillo, 41 km NWCiudad Victoria, 2100 m, Mpo. de Villa Hidalgo(Casas Reales, F14A18)*Small spring W of Coorado Castillo, 41 km NW CiudadVictoria, 2100 m, Mpo. de Villa Hidalgo(Casas Reales, Fl4A18)Sumidero de los Abelos-see Sumidero de Oyamel*Agua de los Allarines, 0.8 km N Coorado Castillo,42 km NW Ciudad Victoria, 1960 m, Mpo. de VillaHidalgo (Casas Reales, Fl4A18)*Cueva de los Allarines, 0.8 km N Comado Castillo,42 km NW Ciudad Victoria, 1900 m, Mpo. de VillaHidalgo (Treacy, 1979:16) (Casas Reales, F14A18)+Cueva de la Boca, 5 km SW Canon de la Boca, 22 kmNNW Ciudad Victoria, Mpo. de ?Guemez (Villa R.,1967:231) (?Guemez, Fl4A19)*Cueva del Borrego, 0.5 km S Comado Castillo, 41 kmNW Ciudad Victoria, 1980 m, Mpo. de Villa Hidago(Casas Reales, Fl4A18)Cueva del Brinco-see Sistema Purificacion*Sotano de las Calenturas, 0.7 km S Yerbabuena,34 km NW Ciudad Victoria, 1460 m, Mpo. de VillaHidalgo (Pate, 1979:86-87) (Casas Reales, Fl4A18)*Cueva de la California, 2 km N Rancho Nuevo,2560 m, Mpo. de Villa Hidalgo (Casas Reales,F14A18)*Cueva del Camino, 1 km W Rancho Nuevo, 34 kmWNW Ciudad Victoria, 2500 m, Mpo. de VillaHidalgo (Casas Reales, F14A18)*Cueva de Coral, near summit of Cerro Zapatero,1 km N Comado Castillo, 41 km NW Ciudad Victoria,2100 m, Mpo. de Villa Hidalgo (Pate, 1979:86)(Casas Reales, F14A18)Cueva de Chuparosa, Rancho Revilla, Mpo. de VillaHidalgo (Casas Reales, Fl4A18)*Cueva de Desmontes, Comado Castillo, 41 km NWCiudad Victoria, 1920 m, Mpo. de Villa Hidalgo(Casas Reales, Fl4A18)+Cueva La Esperanza, 6 km SW Rancho Santa Rosa,32 km NW Ciudad Victoria, 360 m, Mpo. de Guemez(Alvarez, 1963:405) (?Casas Reales, F14A18)*Grieta de las Flores, Conrado Castillo, 41 km NWCiudad Victoria, 2000 m, Mpo. de Villa Hidalgo(Casas Reales, Fl4A18)Cueva de Infiemillo-see Sistema Purificacion*Sotano de Jestis, 1 km S Rancho Revilla, 38 km NWCiudad Victoria, 2210 m, Mpo. de Villa Hidalgo(Casas Reales, Fl4A18)Pozo de Juan Fuentes Perez-see Grieta de las FloresCueva de Musgo, 0.75 km E Rancho Revilla, 38 kmNW Ciudad Victoria, 2100 m, Mpo. de Villa Hidalgo(Casas Reales, F14A18)Cueva del Oso-see Sistema PurificacionCueva de Oyamel-see Sumidero de Oyamel*Sumidero de Oyamel, 1 km N Comado Castillo41 km NW Ciudad Victoria, 1950 m, Mpo. de villiHidalgo (Treacy, 1979:16) (Casas Reales, F14AI8)308
*Sistema Purificacion, Conrado Castillo, 41 km NWCiudad Victoria, 1980 m (Cueva del 080 entrance).1900 m (Cueva del Brinco entrance), 1I00 m(Cueva de Infiemillo entrance), Mpo. de VillaHidalgo (Pate, 1979:82-101, map; Treacy, 1979:8-31,2 maps) (Casas Reales, F14A18)*Sotano de Rancho Nuevo n. 1, Rancho Nuevo,34 km WNW Ciudad Victoria, 2600 m, Mpo. deVilla Hidalgo (Casas Reales, Fl4A18)*Sotano de Rancho Nuevo n. 2, Rancho Nuevo,34 km WNW Ciudad Victoria, 2600 m, Mpo. deVilla Hidalgo (Casas Reales, Fl4A18)*Sotano de Rancho Nuevo n. 3, Rancho Nuevo,34 km WNW Ciudad Victoria, 2600 m, Mpo. deVilla Hidalgo (Casas Reales, F14A18)*Sotano de Rancho Nuevo n. 4, Rancho Nuevo,34 km WNW Ciudad Victoria, 2600 m, Mpo. deVilla Hidalgo (Casas Reales, Fl4A18)Cueva de Rancho Revilla-see Cueva de Revilla*Pozo de Rancho Viejo, 2 km SW Rancho Nuevo,34 km .WNW Ciudad Victoria, 2560 m, Mpo. deVilla Hidalgo (Casas Reales, F14A18)*Cueva de Revilla, Rancho Revilla, 40 km NW CiudadVictoria, 2300 m, Mpo. de Villa Hidalgo (CasasReales, F14A18)Cueva del Tecolote-see Cueva de Oesmontes*Cueva de 105 Tres Bobos, near Conrado Castillo,41 km NW Ciudad Victoria. Mpo. de Villa Hidalgo(Casas Reales, F14A18)Entrada del Viento Alta, Conrado Castillo, 41 kmNW Ciudad Victoria, 2000 m, Mpo. de Villa Hidalgo(Treacy, 1979:17)(Casas Reales, Fl4A18)*Entrada del Viento Baja, Conrado Castillo, 41 kmNW Ciudad Victoria, 2000 m, Mpo. de Villa Hidalgo(Pate, 1979:86; Treacy, 1979:17) (Casas Reales,Fl4A18)Cueva Vrincon-see Sistema Purificacion*Cueva X, Conrado Castillo, 41 km NW Ciudad Victoria,1950 m, Mpo. de Villa Hidalgo (Treacy,1979:16-17) (Casas Reales, F14A18)Santander Jimenez Region+Cueva del Guano, 1I km Nand 21 km W Santanderjimenez, 380 m, Mpo. de Santander jimenez(Aguirre Pequeno, 1959:245-256; Alvarez, 1963:401)Sierra de EI Abra RegionCueva del Abra, 1 km S EI Abra, 9 km NE AntiguoMorelos, 300 m, Mpo. de Antiguo Morelos (Russelland Raines, 1967:65, 67m)(Quintero, Fl4A69)Cueva Chica del Arroyo Seco, 15 km ESE AntiguoMorelos, Mpo. de Antiguo Morelos (Russell,1972c:135)Cueva Grande del Arroyo Seco, 15 km ESE AntiguoMorelos, Mpo. de Antiguo Morelos (Russell,1972c:135)Sotano del Arroyo Seco-see Cueva Grande delArroyo SecoCueva de El Choveno, EI Choveno, 9 km NNE AntiguoMorelos,Mpo. de Antiguo Morelos (Quintero,F14A69)*Cueva de Oiamante, 23 km SE Antiguo Morelos,Mpo. de Antiguo Morelos (Minton, 1978:7-15,map)*Cueva de la Florida, 1 km NNW Praxedis Guerrero(=EI Pachon), 8 km NE Antiguo Morelos, 260 m,Mpo. de Antiguo Morelos (Mitchell, 1970:64m;Russell, 1972c:134-135) (Quintero, Fl4A69)Cueva de EI Mante n. I-see Cueva del Nacimientodel Rio Mante+Cueva del Nacimiento del Rio Mante, 10 km SWCiudad Mante, 100 m, Mpo. de Ciudad Mante(Russell and Raines, 1967:64) (Quintero, F14A69)*Cueva de EI Pachon, Praxedis Guerrero (=EI Pachon),7 km NE Antiguo Morelos, 210.5 m, Mpo. de AntiguoMorelos (Russell and Raines, 1967:65, 68m;Mitchell et aI., 1977:57-58) (Quintero, Fl4A69)Cueva de 105 Pajaros, 1 km NNW Praxedis Guerrero(=EI Pachon), 8 km NE Antiguo Morelos, 260 m,Mpo. de Antiguo Morelos (Russell, 1972c:135)(Quintero, F14A69)*Grutas de Quintero, 1.5 km S Quintero, 13 km SWCiudad Mante, 200 m, Mpo. de Ciudad Mante(Russell and Raines, 1967:64, 66m; Russell,1972c:134) (Quintero, F14A69)Cueva de San Rafael de los Castro8-see Sotano deSan Rafael de los Castros*Sotano de San Rafael de los Castros, San Rafael delos Castros, 13 km ENE Ciudad Mante, 100 m,Mpo. de Ciudad Mante (Russell, 1972c:133)(Lorna Alta, Fl4A59)*Cueva de Santa Elena, 14 km SE Antiguo Morelos,Mpo. de Antiguo Morelos (Russell, 1972c:135136)(Sierra La Colmena, Fl4A79)Sotano de Santa Elena-see Cueva de Santa Elena*Sotano de EI Venadito, 16 km SE Antiguo Morelos,312 m, Mpo. de Antiguo Morelos(Walsh, 1972:12,14m; Mitchell et aI., 1977:56-57 (Sierra La Colmena,Fl4A79)Sierra de Guatemala Region+Cave at Agua Linda, 1I km WNW Gomez Farias,1800 m, Mpo. de Gomez Farias (Martin, 1958:43) (Gomez Farias, F14A49)+Cave 1 km S Aserradero del Paraiso, 13 km NNWChamal, 420 m, Mpo. de Ocampo (Koopman andMartin, 1959:2-3)(Loma Alta, Fl4A59)+Cave, Casa Piedras, 8.5 km WNW Gomez Farias,1500 m, Mpo. de jamauve (Martin, 1958:44)(Gomez Farias, F14A49)+Cave, Ojo de Agua, 3 km SE Gomez Farias, 100 m,Mpo. de Gomez Farias (Alvarez, 1963:398)(Gomez Farias, F14A49)+Caves, Rancho del Cielo, 7 km NNW Gomez Farias,1I60 m, Mpo. de Gomez Farias (Hooper, 1953:9) (Gomez Farias, Fl4A49)+Bell-shaped sink, Rancho del Cielo, 7 km NNWGomez Farias, 1I60 m, Mpo. de Gomez Farias(Martin, 1958:47) (Gomez Farias, F14A49)+Ory sinkhole, Rancho del Cielo, 7 km NNW GomezFarias, 1I60 m, Mpo. de Gomez Farias (Martin,1958:56) (Gomez Farias, Fl4A49)+Deep open sink, Rancho del Cielo, 7 km NNW GomezFarias, 1I60 m, Mpo. de Gomez Farias (Martin,1958:43) (Gomez Farias, Fl4A49)+Sinkhole, 12 m deep (No.1), Rancho del Cielo, 7 kmNNW Gomez Farias, 1I60 m, Mpo. de Gomez Farias(Martin, 1958:47) (Gomez Farias, Fl4A49)309
Page 1:
A REVIEW OF THECAVERNICOLE FAUNA OF
Page 8:
. 1-----,, ,", ,\'"I/I~" )/Fig. I.-
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collected a troglobitic cirolanid i
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C. Bolivar and J. Carranza investig
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springs in the vicinity of Cautro C
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Cueva del Diablo near Orizaba, Vera
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Those species of troglobite which h
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Table 2.-(continued)1970. Eodrilus
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CAVE REGIONSOnly the cave regions o
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Table 3.-(continued)TroglobitesTrog
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y the waters of the Amistad Reservo
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Although a few caves in the Norther
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an elevation of about 900 meters. T
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the gryllid cricket Paracophus suba
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has been Jiscussed by Segerstrom (1
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only apparent troglophile is the ps
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The cave fauna of this region is ri
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igneous rock immediately east of th
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Rio Soto la Marina and terminates a
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caves studied are many large stream
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unusual species is Neogovea mexasca
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pallidus Rowland, the nesticid spid
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troglobites: the trichoniscid isopo
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Cambridge, and Pholcophora bispinos
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of the genus Lutzomyia has not been
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The principal cavernous fonnation i
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ocks of Paleocene to Pliocene age.
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Beach and Supra-tidal Zone, the I\o
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The aquatic fauna of Mexico and Cen
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oaxacae Hobbs from the Valle Nacion
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described, the genus occurs in cave
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large genus with numerous epigean s
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milliped, laniatorid harvestmen, an
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Table 4.-Summary of cave inhabiting
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Discussion.-This large, troglobitic
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any way a part of the true cave fau
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they have been washed in with soil
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part of the aquatic fauna. A very l
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The genus Helicodiscus is a signifi
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associated with true cave waters. P
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Family DaphnidaeWilson (1936) repor
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tul), Cueva de San Isidro, Cueva de
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30 m below entrance of Cueva de la
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underflow of rivers, alluvial grave
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the similarities between the distri
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Yucatan but did not identify the sp
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Protrichoniscus: Reddell, 1967a:24.
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It indudes five spccies, of which t
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Type-locality.-Pozzo, Etla, Oaxaca,
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Records.-Campeche: Volc
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164; Pearse, 1945:167, 168, fig. 13
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Family AtyidaeThe family Atyidae is
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Creaseria morleyi (Creaser)Blind sh
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the pond which is fed by the cave s
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philes. Pseudothelphusa (Pseudothel
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Typhlochactas elliotti MitchellTyph
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Distribution.-Known only from the t
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(Dumitresco and Orghidan, 1977); th
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Leucohya magnifica MuchmoreUndescri
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Discussion.-This is the largest spe
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Type-Iocality.-S6tano de Huitzmolot
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Bartolo is a name applied to two ne
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Rowland, 1975b:19, 20; Dumitresco,
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Type-Iocality.-Actlin Kaua, Yucatan
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Distribution.-Known only from caves
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een collected. It is frequently fou
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Distribution.-Known from two caves
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actually a natural cave artificiall
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Discussion.-This large species has
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and T. patellaris Bryant from caves
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estricted to the Mexican Plateau (s
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maya Chamberlin and Ivie from Campe
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Metagonia chiquita GertschMetagonia
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Metagonia capilla: Mitchell and Kaw
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the cave and in general association
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DiscWlSion.-This is a delicate, eye
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Type-Iocality.-Cueva de Los Sabinos
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family. Clarence and Marie Goodnigh
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n. 2, San Luis Potosi; and K. rugos
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Type-locality.-Cueva de Apoala, San
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Records.-Chiapas: Sumidero de Canad
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Family HeterozerconidaeSpecimens of
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only one species. All known epigean
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nation must await study, but it is
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Mexico and Guatemala, and several u
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Type-locality.-Gruta de Atoyac, Ver
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Records.-San Luis PotOSI: Sotano de
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Records.-Veracruz: Grutas de Atoyac
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the genus Synoptura and removed the
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known from caves all contain troglo
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las Perlas, Cueva Chica de la Perra
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Discussion.-This species is distinc
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Speodesmus pecki ShearPolydesmidae,
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appears to be most closely related
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this is almost certainly M. russell
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Table 21.-Summary of cave inhabitin
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Type-Iocality.-Cueva Bonita del Pre
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Type-Iocality.-Cueva de El Jobo, El
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Order DipluraFamily CampodeidaeAlth
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of small soil inhabiting japygoids
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(Linnaeus) has been collected from
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Cave. Paracophus caecus appears to
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the most poorly known species in th
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Family EpipsocidaeThe family Epipso
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citata Van Tyne in Cenote de Piste,
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een found in caves and mines in Yuc
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ground dwelling habit, many beetles
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Type-Iocality.-Sotano de Tejamanil,
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Vigna Taglianti, 1972:118, 121-127,
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Veracruz) belongs in another subgen
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Troglobacanius sbordonii VomeroTrog
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Ptomaphagus (Adelops) is a group of
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(A.) speLaeus, from Grutas de Cacah
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epresented by many species. Tenebri
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Myodopsylla gentilis Jordan and Rot
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genus Pholeomyia have been collecte
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Wheeler (1938) has reported the pre
Page 246 and 247:
263.264, 265, 267, 270, 273, fig. 5
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Anophthyctys jordani: Stefanelli, 1
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Prietella phreatophila Carranza, 19
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TyphlilJs: Pearse, 1938a:13, 15; Hu
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Family BufonidaeFour species of toa
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Oaxaca; and E. yucatanensis Lynch f
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sity of bats in the tropics. In add
Page 260 and 261:
Three species of short-tailed bats
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abundant in the entrance area of Cu
Page 264 and 265: Andrews, E. W., IV. 1970. Balankanc
Page 266 and 267: Bolivar y Pieltain, C. 1940. Explor
Page 268 and 269: Brignoli, P. M. 1979a. The morpholo
Page 270 and 271: Conde, B. 1975. Description du prem
Page 272 and 273: Ediger, G. 1970. The Birthday Passa
Page 274 and 275: Gonzalez Ochoa, A. 1963b. Relacione
Page 276 and 277: Hobbs, H. H.,Jr., and T. C. Barr, J
Page 278 and 279: Jordan, c. B. 1937. Bringing in the
Page 280 and 281: McKenzie, D. 1965a. Caves of the Si
Page 282 and 283: Monod, T. 1975. Sur la distribution
Page 284 and 285: Peck, S. B., and J. J. Lewis. 1977.
Page 286 and 287: Reddell, J. R., and W. R. Elliott.
Page 288 and 289: Sanborn, C. C. 1936. Records and me
Page 290 and 291: Stone, B. 1979. The 1979 San Agusti
Page 292 and 293: Villa R., B. 1956. Una extrana y se
Page 294 and 295: Wimsatt, W. A. 1969. Transient beha
Page 296 and 297: +*Cueva de los Chivos, 5 km E Altam
Page 298 and 299: Sumidero Yochib, Yochib, 8 km NE Te
Page 300 and 301: Serrania del Burro RegionCave, Haci
Page 302 and 303: +Cueva de Tezoapa, 1.5 km E Acahuiz
Page 304 and 305: *Gruta Sur de San Bartolo, 9 km S S
Page 306 and 307: *Cueva del Puente de Fierro, 1.5 km
Page 308: Cueva de los Riscos, 2 km S Rancho
Page 311 and 312: Cueva de El Coy-see Cueva del Nacun
Page 313: +*Cueva del Aire, 1.8 km at S85°W
Page 317 and 318: *Cueva del Nacimiento del Rio Frio,
Page 319 and 320: Sotano de Humo, 5 km NW San Andres,
Page 321 and 322: and Bohor, 1975:25-27, 26m; Reddell
Page 323 and 324: *Actun Xpukil, 3 km S Calcehtok, Mp
Page 325 and 326: Appendix 2LIST OF TROGLOBITES BY ST
Page 327 and 328: Paravachonium superbum-I'seudoscorp
Page 329 and 330: SUPPLEMENTWhile this manuscript was
Page 331 and 332: Cambala speobia (Chamberlin) (p. 19
Page 333: mida (Arachnida) in the New World.
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