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searchable PDF - Association for Mexican Cave Studies

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appears to be most closely related to Ergene setosusChamberlin, an epigean species known from San LuisPotosi and Tamaulipas. Other typhlobollelids includetwo species of Typhlobolellus from Veracruz andpossibly Morelene from Morelos. Reddellobus troglobiuswas found in areas of organic debris and batguano in each of the recorded caves. Although alltypWobollelids lack eyes, this species has reducedpigmentation, slender body, and attenuate appendages,indicating that it should be considered to be atroglobite.Family SpirobolidaeTwo species of this family have been collectedfrom caves in Mexico. Hiltonius carpinus carpinusChamberlin was found in the entrance area of Grutasdel Mogote and Cueva Chica del Mogote, Guerrero.An undetermined genus and species has been collectedin a single cave in Nuevo Leon.Order SpirostreptidaFamily CambalidaeThe family Cambalidae is represented in the cavesof Mexico and Belize only by troglobitic species. Thisfamily is well represented in the caves of the southernUnited States but also abounds in <strong>for</strong>ested areas. Nocollections of the family from the epigean environmentare known <strong>for</strong> Mexico, Guatemala, or Belize.Cambala speobia (Chamberlin)Eclytus (not Holmgren 1855) speobius Chamberlin,1952b:1O.Eclomus speobius: Chamberlin, 1952a:71; Chamberlinand Hoffman, 1958:175.Cambala caeca Loomis, 1953:417, fig. 1-3.Cambala captiosa Causey, 1959:67-71, fig. 1-3.Cambala speobia: Causey, 1964a:243-244; Causey,1971a:272, 273, fig. la, 2a; Shear, 1974:249.Cambala speobia new subspecies: Reddell, 1965b:164-165; Loomis, 1968:95.Cambala speobia subspecies: Causey, 1965:63; Mitchelland Reddell, 1971:64; Reddell, 1971b:46.Cambala: Causey, 1968:124.Type-Iocality.-Of Eclytus speobius: Wyatt <strong>Cave</strong>,Sutton (=Edwards) County, Texas, U.S.A.; of Cambalacaeca: Wyatt <strong>Cave</strong>, Edwards County, Texas,U.S.A.; of Cambala captiosa: Beck's Ranch <strong>Cave</strong>,Williamson County, Texas, U.S.A.Distribution.-Known from many caves in CentralTexas (U.S.A.) and one cave in northern Coahuila,Mexico. See Fig. 62.Mexico record.-Coahuila: Cueva de los Lagos.Discussion.-This is the common cave milliped ofCentral Texas. Its occurrence in a cave 2 km south ofthe Rio Grande in Coahuila is not surprising. Thelimestones of ValVerde County, Texas, are identicalto those in adjacent Coahuila. Specimens from Coahuilaappear to exhibit slight differences from thoseacross the Rio Grande, and, at first, these appearedto be sufficient to justify subspecific recognition <strong>for</strong>the <strong>Mexican</strong> population. Careful study of all materialfrom Central Texas has shown that these differencesfall within the range of variation within the speciesin Texas. The species is closely related to Cambalareddelli reddelli Causey in northwestern Texas andadjacent New Mexico and C. reddelli inomatusCausey in northwest Texas. Both of these <strong>for</strong>ms arealso cavernicolous. Cambala speobia was collectedfrom silt and guano in a short horizontal passage inthe upper levels of Cueva de los Lagos. This cave hasnow been inundated by the waters of the AmistadReservoir.Jarmilka alba ShearJarmilka alba Shear, 1973:44-45, fig. 1-3.Type-locality -Mountain Cow <strong>Cave</strong>, <strong>Cave</strong>s Branch,Belize.Distribution.-Known only from the type-locality.See Fig. 62.Discussion.-The ecological status of this <strong>for</strong>m is indoubt, and Shear (1973) expresses the opinion that itmay be a facultative troglophile also occurring in theendogean habitat. It is included here because of itslack of eyes and pigment and because all known<strong>Mexican</strong> cambalids appear to be restricted to caves.This is the smallest species in the family and the mostsouthern representative of the Cambalidae in NorthAmerica.Mexicambala blanda CauseyMexicambala russelli undescribed subspecies: Reddell,1966a:13; Reddell, 1967c:55 (all records exceptSotano de la Joya de Salas and Cueva de los Leones);Reddell, 1971b:47 (all records except Sotanode la J oya de Salas and Cueva de los Leones).Mexicambala sp.: Reddell and Mitchell, 1971b:190(all records except Cueva de la Capilla, Sotano dela Joya de Salas, and Cueva de los Leones).Mexicambala blandus Causey, 1971a:272, 275, 276,278-279, fig. lc, 2b, 3; Reddell, 1973a:35, 40.Mexicambala: Reddell, 1973b:77.Mexicambala blanda: Causey, 1973:121; Reddell andElliott, 1973b:182, 185; Shear, 1974:250.Mexicambala russelli: Vomero, 1974:350 (Sotano deGomez Farias record only).192

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