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Vergara - 1976 - Physiological and morphological adaptability of ri

Vergara - 1976 - Physiological and morphological adaptability of ri

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GENETIC VstRlOLlSNESS IN CLIAIAIIC ADAPTATION I09<br />

water temperature under different depths on the growth. grain yield <strong>and</strong> yield-components <strong>of</strong><br />

<strong>ri</strong>ce [in Japanese, English sununarfl. Proc. Crop Sci. Soc. Jpn. 31:19-22.<br />

VERGARA. B. S., T. T. CHANG, <strong>and</strong> R. LILIS. I972. The flowe<strong>ri</strong>ng response <strong>of</strong> the <strong>ri</strong>ce plant to<br />

photope<strong>ri</strong>od. Int. Rice Res. Inst. Tech. Bull. Szl-SS (re\-'.).<br />

VERGAILA. B. S.. T. M. CHU. <strong>and</strong> R. M. VISPERAS. 1970. EtTect <strong>of</strong> temperature on the anthesis<br />

<strong>of</strong> IRS. Int. Rice Comm. Newsl. l9(3):ll-l7.<br />

\r'ERG.-\RA. B. S.. R. LtLIS, A. BUENO, <strong>and</strong> R. DONATO. 1967. Adaptability to cultural conditions<br />

<strong>of</strong> Raminad Strain 3. Philipp. Ag<strong>ri</strong>c. 51:253-267.<br />

VERGARBI. B. S.. S. PURAN.-\Bl-LAVUNG. <strong>and</strong> R. LILIS. 1965. Factors determining the growth duration<br />

<strong>of</strong> <strong>ri</strong>ce va<strong>ri</strong>eties. Phyton 22:l77-1S5.<br />

“IADA. E. I952. Studies on the response <strong>of</strong> heading to daylength <strong>and</strong> temperature in <strong>ri</strong>ce plants.<br />

I. Response <strong>of</strong> va<strong>ri</strong>eties <strong>and</strong> the relation to their geographic dist<strong>ri</strong>bution in Japan [in Japanese.<br />

English summaryi]. Jpn. J. Breed. 2:55-62.<br />

WADA, E. I954. Studies on the response <strong>of</strong> heading for day~length <strong>and</strong> temperature in <strong>ri</strong>ce<br />

plants. II. Response <strong>of</strong> upl<strong>and</strong> <strong>ri</strong>ce <strong>and</strong> foreign <strong>ri</strong>ce <strong>and</strong> the relation to their geographical dist<strong>ri</strong>bution<br />

[in Japanese. English summaryi]. Jpn. J. Breed. 3:22-26.<br />

WU. H. P. 1968. Studies on the quantitative inhe<strong>ri</strong>tance <strong>of</strong> (Jljira satwa L. l. A diallcl analysis<br />

<strong>of</strong> heading time <strong>and</strong> plant height in F, progeny. Bot. Bull. Acad. Sin. 9:I~9.<br />

YAO. Y. T“ <strong>and</strong> C. J. YU. I963. Biomet<strong>ri</strong>sche Studien uher die Vererbung der z-‘tusschusszeiten<br />

Beim Reis. Bot. Bull. Acad. Sin. 41140-162.<br />

Yosttlmc S., J. H. COCK. <strong>and</strong> F. T. Pacino. 1972. <strong>Physiological</strong> aspects <strong>of</strong> high yields. Pages<br />

454-469 m Intemational Rice Research Institute, Rice breeding. Los Banos. Philippines.<br />

YU. C. J.. <strong>and</strong> Y. T. YAo. 1957. Uber die Vererbung der Ausschusszeiten Beim Reis. Jpn. J.<br />

Genet. 32:179-188.<br />

DISCUSSION<br />

STANSEL (Chairman): How many here while reading through the literature have come across<br />

passages such as. “Observations were incomplete due to high temperature“? Or this: “Low<br />

temperatures <strong>and</strong> excessive rainfall may have influenced the results." with no definition <strong>of</strong> these<br />

climatic factors. How many times do we see these references to climatic va<strong>ri</strong>ables in the literature<br />

without definition? Everyone recognizes the impact <strong>of</strong> climatic va<strong>ri</strong>ables but too few report<br />

them. This is the problem our participants faced in accumulating the reported observations<br />

from the literature. Have you been guilty <strong>of</strong> this practice?<br />

Breeding for the illusive climatic <strong>adaptability</strong> can be a hazardous endeavor. Gains in some<br />

characte<strong>ri</strong>stics can <strong>of</strong>ien be at the det<strong>ri</strong>ment <strong>of</strong> other characters. How <strong>of</strong>ten does our search for<br />

climatic <strong>adaptability</strong> result in a narrowing <strong>of</strong> climatic response in the plant? For example, we<br />

discussed earlier the development <strong>of</strong> US. va<strong>ri</strong>eties which were adapted to increased utilization<br />

<strong>of</strong> available sunlight. resulting in increased yield. Yes, yields have been significantly increased.<br />

but for a p<strong>ri</strong>ce. These very early matu<strong>ri</strong>ng va<strong>ri</strong>eties also require a very high level <strong>of</strong> cultural<br />

control. Water management <strong>of</strong> these va<strong>ri</strong>eties is c<strong>ri</strong>tical due to the short basic vegetative phase.<br />

Both timing <strong>and</strong> rate <strong>of</strong> fertilizer is c<strong>ri</strong>tical for high yields. Weed control has caused producer<br />

problems due to inability to compete with moderate weed infestations. Yields are so dependent<br />

on climatic conditions. These va<strong>ri</strong>eties then have specific or limited climatic <strong>adaptability</strong>.<br />

While they do well in Texas. they are not adapted elsewhere. How <strong>of</strong>ten do you hear. “We<br />

have developed resistance to a particular disease; however. yields under disease-tree conditions<br />

do not match the yields <strong>of</strong> local va<strong>ri</strong>eties"? Yes, too <strong>of</strong>ten we arc etfcctive in developing yield<br />

resistance because <strong>of</strong> a rest<strong>ri</strong>ctive climatic <strong>adaptability</strong>‘. ls the climate in your locality the same<br />

year afier year‘? How <strong>of</strong>ten do you Iiear. “There is no such thing as a normal year"? Normality<br />

is the exception to the rule. Plant breeders, more than any other group. are constantly trying to<br />

measure climatic response. The program here at IRRI has shown there are eFfective ways to<br />

develop climatic <strong>adaptability</strong>.<br />

I wish to reiterate the observations discussed earlier by Dr. Chang because there is a message<br />

here for all stibject-matter areas <strong>of</strong> ag<strong>ri</strong>cultural research. Disruptive seasonal selection is

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