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Vergara - 1976 - Physiological and morphological adaptability of ri

Vergara - 1976 - Physiological and morphological adaptability of ri

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'l'El\‘lPERA'l'LlRE AND LIGHT. REPRODUCTTVE GROVWTH AND RLPENING T99<br />

0 9<br />

O8 —<br />

0 7 —<br />

O6 —<br />

Heading<br />

Q 1 I | l l | | I | |<br />

-40 _2o O +20 + 4O<br />

Stage from heouing tnrrvetdcys)<br />

9. Change in multiple correlation coefiicient <strong>of</strong> <strong>ri</strong>pening<br />

gade ‘with grouth stage. Model: Modified type IV<br />

(ltlj) - 11,715‘ l b_TS » b.~TS - lnTS I b,1" It bu?<br />

+ b1T+ In T + c<br />

where T = l0‘ exp (~ 7.000 {I l 273). 2)<br />

S = s i 4.5<br />

t <strong>and</strong> s: mean values for 20 days <strong>of</strong> temperature <strong>and</strong><br />

sunshine hours.<br />

The trough between two peaks is cqual to heading timc. The fact that thc<br />

effect <strong>of</strong> climate at heading time is relatively small is interesting. This may be<br />

partially" att<strong>ri</strong>buted to small va<strong>ri</strong>ance <strong>of</strong> climatic factors at heading time. The<br />

st<strong>and</strong>ard deviation <strong>of</strong> temperature throughout the whole growing stage becomes<br />

minimal near heading time (sec Fig. 2). This tendency was not observed xvhen<br />

the data were divided into several groups ‘with similar temperature conditions.<br />

However the declines in multiple correlation coefficient (R) at heading were still<br />

observed in each <strong>of</strong> the groups.<br />

Thus. it appears that this phenomenon is not necessa<strong>ri</strong>ly caused by the small<br />

va<strong>ri</strong>ance <strong>of</strong> temperature.<br />

Shibata ct al. (1970) found that the occurrence <strong>of</strong> ste<strong>ri</strong>le paddy by low" temperature<br />

treatment was more se<strong>ri</strong>ous at the spikelet differentiation stage than at<br />

heading time. It was also pointed out that when the pe<strong>ri</strong>od <strong>of</strong> low temperature<br />

treatment was not so long, the <strong>ri</strong>ce plant might escape chill injury b)" delaying<br />

the heading time.<br />

lt was difficult to explain, however, why the decline in R [sensitivity to climate)<br />

at heading was observed even in warm regions.<br />

Functional relationships between <strong>ri</strong>pening grade <strong>and</strong> temperature<br />

When sunshine hour (s) in eq. (2) is fixed. the regression curve <strong>of</strong> the <strong>ri</strong>pening<br />

grade on temperature at four stages is shown in Fig. l0. From this. it is found<br />

that temperature curves <strong>of</strong> <strong>ri</strong>pening grade differ with growth stages.<br />

At l- 20] stage. which is one <strong>of</strong> two stages xvith higher R value. the optimum<br />

temperature for <strong>ri</strong>pening grade is about 23°C (Fig. 10A). The characte<strong>ri</strong>stic <strong>of</strong>

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