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Vergara - 1976 - Physiological and morphological adaptability of ri

Vergara - 1976 - Physiological and morphological adaptability of ri

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RICE GROWTH l.\l DII~'FERE.\IT' ENVIRONIVIEN'I‘S 435<br />

Type III: A slow growth at early stages followed by a high growth rate until<br />

grovrth ceases; frequently observed in high-latitude areas where the temperature<br />

is low in sp<strong>ri</strong>ng; generally‘ the total dry-matter production is small, <strong>and</strong> sometimes<br />

the grain yield is moderate due to a reasonable grain-straw ratio but<br />

frequently the grain yield is low due to a small grain-straw ratio.<br />

Type IV: Continuous slow growth; caused by insufficient nut<strong>ri</strong>tional supplies.<br />

adverse soil conditions, ete.; very low grain yields; this tyjpe is not discussed<br />

in this paper because climatic environment is not the growth-limiting<br />

factor.<br />

Type V (V'): A moderate (or high) growth rate followed by a high growth rate<br />

du<strong>ri</strong>ng later stages; possible only’ with abundant solar radiation du<strong>ri</strong>ng <strong>ri</strong>pening<br />

(or throughout the growth) with good plant type va<strong>ri</strong>eties; an extremely high<br />

grain yield due to large values <strong>of</strong> the total dry-matter production as well as the<br />

grain-straw ratio.<br />

In a va<strong>ri</strong>etal compa<strong>ri</strong>son <strong>of</strong> grow-th. Oka et al. (1970) desc<strong>ri</strong>bed early <strong>and</strong><br />

sustained vigor types. These correspond to Type I <strong>and</strong> Type II. respectively‘.<br />

lf we look at va<strong>ri</strong>ous types <strong>of</strong> growth patterns from the st<strong>and</strong>point <strong>of</strong> phasal<br />

development the following three types can be classified (Fig. 3) (Tanaka et al._<br />

1964).<br />

Type A (vegetative phase—rcproduetive phase): The paniele-p<strong>ri</strong>mordia are<br />

initiated soon after the maximum tiller number stage; considered to be normal;<br />

the growth curve is generally Ty-"pe II or Type V; uniform flowe<strong>ri</strong>ng; generally<br />

the grain yield is high.<br />

'l"_vpe B (vegetative phase—reproductive phase overlapped with vegetative<br />

phase) : Tille<strong>ri</strong>ng continues after the paniele-p<strong>ri</strong>mordia initiation stage‘, generally‘<br />

caused by a continuous active nitrogen uptake which is induced by a slow nitrogen<br />

uptake at early growth stages due to low temperatures, excess nitrogen application.<br />

untimely nitrogen top-dressing. excessively’ wide spacing‘ use <strong>of</strong> unduly<br />

short-duration va<strong>ri</strong>eties etex. the growth curve is irften Type lll; flowe<strong>ri</strong>ng is<br />

irregular due to a wide age va<strong>ri</strong>ation among tillers; the number <strong>of</strong> spikelets per<br />

panicle is small because the photosynlhale after the panicle-p<strong>ri</strong>mordia initiation<br />

stage is consumed not only by the panicle-p<strong>ri</strong>mordia but also by the new-"lf;<br />

developing tillers; frequently associated with low grain yields.<br />

Type C (vegetative phase—vegetative-lag phase—reproductive phase) :<br />

Tille<strong>ri</strong>ng<br />

diseontinues long before the panicle-p<strong>ri</strong>mordia initiation stage; occurs when<br />

the vegetative phase is extremely long; associated with high temperatures from<br />

transplanting. <strong>and</strong> exhaustion <strong>of</strong> nitrogen supply in the soil from an early growth<br />

stage; the growth curve is generally Type l; the number <strong>of</strong> panicles per unit field<br />

area tends to be small due to a high percentage <strong>of</strong> ineffective tillers; the number<br />

<strong>of</strong> spikelets is frequently small due to insufficient supply <strong>of</strong> nitrogen du<strong>ri</strong>ng the<br />

reproductive phase; the grain yield is <strong>of</strong>ten low. As the dry-matter production is<br />

slow du<strong>ri</strong>ng the pe<strong>ri</strong>od between the maximum tiller number stage <strong>and</strong> the panicle-p<strong>ri</strong>mordia<br />

initiation stage, this pe<strong>ri</strong>od is called the vegetative-lag phase.<br />

Du<strong>ri</strong>ng this phase the internodes at the base start to elongate before the paniele-

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