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Vergara - 1976 - Physiological and morphological adaptability of ri

Vergara - 1976 - Physiological and morphological adaptability of ri

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202 crux-tars AND RICE<br />

At such low mean temperatures. <strong>ri</strong>ce plants in the field may have many chances<br />

<strong>of</strong> chill injury <strong>and</strong> frost, because <strong>of</strong> daily va<strong>ri</strong>ation in temperature <strong>and</strong> rapid fall<br />

<strong>of</strong> night temperature. This seems to explain why the optimum temperature for<br />

<strong>ri</strong>pening is higher in the field than in the pliyttatrtin.<br />

The time course 0f optimum temperature can be obtained from the temperature—<strong>ri</strong>pening<br />

grade curves in each stage (Fig. 10). With 6 hours <strong>of</strong> sunshine.<br />

the optimum temperatures are 230° at 1-20], 26.0°C at heading. <strong>and</strong> 225°C<br />

at [ +20] stage.<br />

Relationships between light <strong>and</strong> <strong>ri</strong>pening grade<br />

The effect <strong>of</strong> light on <strong>ri</strong>pening grade may vary depending upon stage <strong>and</strong><br />

temperature. Figure 11 indicates that the effect <strong>of</strong> sunshine hours is positive<br />

under high temperature conditions. irrespective <strong>of</strong> stage, while negligible or even<br />

negative under low temperature conditions, especially at [+ 10] stage. Wada et<br />

a1. (1972) reported that the effect <strong>of</strong> light intensity on the <strong>ri</strong>pening grade (percentage<br />

<strong>of</strong> ste<strong>ri</strong>le grains) W35 negligible at 18°C du<strong>ri</strong>ng the meiosis stage. Similar<br />

results were obtained by Kiyosawa (1962). However. there is no report about<br />

the negative effect <strong>of</strong> light on <strong>ri</strong>pening under low temperature. Negative effect <strong>of</strong><br />

sunshine hours under low temperature conditions may be att<strong>ri</strong>butable to low<br />

night temperatures associated with long sunshine hours. The correlation between<br />

sunshine hours <strong>and</strong> diurnal temperature was 0.74 (p < 0.1%) in Kitami in the<br />

cool region.<br />

Munakata (1968) ObSBTVCd that <strong>ri</strong>pening grade <strong>of</strong> the <strong>ri</strong>ce plant tvas decreased<br />

by strong solar radiation (over 450 cal -em-= - day-l at high temperature above<br />

27°C). Such a tendency was scarcely found in these statistical data.<br />

Estimation 0f <strong>ri</strong>pening grade<br />

The accuracy <strong>of</strong> estimation <strong>of</strong> <strong>ri</strong>pening grade was considered to be increased<br />

by the equation which included climatic data at three stages. i.e. at [-20]<br />

(spikelet differentiation stage), [0] (heading stage) <strong>and</strong> [— 20] (<strong>ri</strong>pening stage).<br />

The model used was the type IV modified eq. (2) (type III, including six terms).<br />

In this model, five terms were used for each stage <strong>and</strong> the total was 15, because<br />

<strong>of</strong> limitation <strong>of</strong> memory size in our mid-computer. The regression formula <strong>of</strong><br />

<strong>ri</strong>pening grade was as follows:<br />

(I/Yaa: = _ — 4.33 - 101. T,"~-‘-.S';= + 2.34-10-1-T,'1 - S,"<br />

+ 6.97-101-T,°-S‘;1 + 1.80 - 10" T5“ S,“<br />

+ 1.33-10**T;*-‘ S,“ + 1.75-101T+* S‘<br />

— 1.30 - 10 ' TJS,“ - 1.84- 10 7982-1 — 6.08 - 10-’ T,“ 5'9“<br />

— 1.88-10‘7',-’.S',°-“ + 1.47 ~ 101 7'," 5}“<br />

— 4.27 - 10’ T98," + 2.34 - 10 =T;' S," + 1.8716 (3)<br />

where T,<br />

<strong>and</strong><br />

I}; <strong>ri</strong>pening grade (Ti/Y; -mg).

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