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Vergara - 1976 - Physiological and morphological adaptability of ri

Vergara - 1976 - Physiological and morphological adaptability of ri

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as a function <strong>of</strong> solar elevation in Fig. 2.<br />

IMHCROCLISLATE or THE RICE CROP 119<br />

The decrease <strong>of</strong> value <strong>of</strong> lo, with increasing<br />

solar elevation implies that direct solar radiation can penetrate deeper<br />

when the solar elevation is higher. The dependence <strong>of</strong> k, on solar elevation<br />

presented in Fig. 2 resembled that for a model canopy" consisting <strong>of</strong> leaves with<br />

an inclination <strong>of</strong>70° to 75° (eg. Isobe. 1969).<br />

Diffuse radiation<br />

As already desc<strong>ri</strong>bed, diffuse radiation flux in plant canopies consists <strong>of</strong> two<br />

components <strong>of</strong> diffuse solar radiation tSDR) <strong>and</strong> complementary diffuse radiation<br />

due to scatte<strong>ri</strong>ng <strong>of</strong> light by plant elements (CDR). It is reasonable to<br />

assume that the extinction <strong>of</strong> both diffuse radiation fluxes in plant canopies is<br />

given by Beefs law. The intensity <strong>of</strong> SDR penetrating to a leaf area index <strong>of</strong> L<br />

is desc<strong>ri</strong>bed by<br />

ML) = $.40) EXPFRAL) (5)<br />

where S10) is the SDR intensity at the top <strong>of</strong> plant cantipy, <strong>and</strong> k, is the extinction<br />

coefficient for diffuse radiation. On tl1e basis <strong>of</strong> research by Monsi <strong>and</strong><br />

Saeki (i953). the following values <strong>of</strong> kd were determined:<br />

Manryo canopy k,, = 0.67-0.70‘<br />

[R8 canopy k,, = 0.60<br />

The hatched area in Fig. 3 denotes vertical pr<strong>of</strong>iles <strong>of</strong> CDR emanating from<br />

each unit leaf lajver (B(z) <strong>and</strong> C(z)). As shown in Fig. 3. when solar elevation is<br />

lovi“. the flux <strong>of</strong> CDR emanating from each unit leaf layer is not large <strong>and</strong> its<br />

peak value is about '7 vvlml. With increasing solar elevation, the generation <strong>of</strong><br />

CDR is observed throughout the whole foliage layer <strong>of</strong> the <strong>ri</strong>ce crop. mainly<br />

because <strong>of</strong> deeper penetration <strong>of</strong> direct solar radiation. The leaf area depth at<br />

tvhich the CDR was maximum moved gradually dowmvard with increasing solar<br />

elevation <strong>and</strong> reached 0.6-0.7 <strong>of</strong> canopy} height. Pltnvever, the depth at which<br />

CDR was maximal was consistently higher than the layer with the maximum <strong>of</strong><br />

leaf area density (LAD)<br />

The upward flux <strong>of</strong> CDR decreased monotonically downward. The upward<br />

flux <strong>of</strong> CDR at the top <strong>of</strong> crop canopy characte<strong>ri</strong>zing the radiation flux reflected<br />

from <strong>ri</strong>ce canopies increased with increments <strong>of</strong> solar elevation. The pr<strong>of</strong>iles <strong>of</strong><br />

doo/nn-"ard flux <strong>of</strong> CDR are characte<strong>ri</strong>zed by the curve with a rapid increase in<br />

the upper half layer <strong>of</strong> the canopy <strong>and</strong> a slower decrease in the bottom cantipv<br />

layer. The slower decrease <strong>of</strong> downward flux <strong>of</strong> CDR in the bottom half layer<br />

<strong>of</strong> the canopy was mainly because <strong>of</strong> the steep decrease <strong>of</strong> leaf area density in<br />

this region.<br />

Radiation intensity on leaf surfaces<br />

The radiant energy impinging on leaf surfaces within a plant canopy is known<br />

to play a decisive role in plant evaporation <strong>and</strong> canopy photosynthesis. The<br />

irradiance <strong>of</strong>leafsurfaces (S) can be expressed as

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