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Vergara - 1976 - Physiological and morphological adaptability of ri

Vergara - 1976 - Physiological and morphological adaptability of ri

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RICE GROWTH IN DIFFERENT Emotions-isms 437<br />

GROWTH PATTERNS IN DIFFERENT ENVIRONMENTS<br />

Figure 4 illustrates the diversity <strong>of</strong> locations <strong>and</strong> seasons <strong>of</strong> <strong>ri</strong>ce cultivation in<br />

Asian count<strong>ri</strong>es (after P. Kung. mimeo). The environment <strong>of</strong> cultivated <strong>ri</strong>ce<br />

va<strong>ri</strong>es from place to place. from season to season at a given place. <strong>and</strong> also from<br />

year to year at a given place <strong>and</strong> season. Although selection <strong>of</strong> va<strong>ri</strong>eties is made<br />

by conside<strong>ri</strong>ng the environmental condition in which the crop is going to be<br />

grown, the growth pattern va<strong>ri</strong>es considerably. Thus. va<strong>ri</strong>ous cultural practices<br />

are used to manipulate the growth pattern to obtain maximum grain yield. In<br />

this sense the growth pattem is the result <strong>of</strong> environment-va<strong>ri</strong>ety-cultural<br />

practice interaction.<br />

Locations<br />

Long-duration va<strong>ri</strong>eties cannot be grown in high-latitude areas because<br />

<strong>of</strong> the limited season with suitable temperatures. but they are popular in<br />

low-latitude areas for va<strong>ri</strong>ous reasons. Commercial medium-duration va<strong>ri</strong>eties,<br />

however. mature in about 130 days throughout <strong>ri</strong>ce-grouting areas. In highlatitude<br />

areas the vegetative growth is slow <strong>and</strong> long. whereas in towlatitude<br />

areas it is vigorous <strong>and</strong> short. Because <strong>of</strong> this. there are more chances for<br />

medium-duration va<strong>ri</strong>eties to assume Type B<strong>and</strong> Type C patterns in high <strong>and</strong><br />

in low-latitude areas, respectively’.<br />

With medium-duration va<strong>ri</strong>eties the time from stiwing to floive<strong>ri</strong>ng is almost<br />

constant, but the lealing interval is shorter at higher temperatures. Thus. the<br />

number <strong>of</strong> leaves on the main stem is greater in loxv= than in high-latitude areas.<br />

The three leaves from the flag leaf on each stem are the ones which cont<strong>ri</strong>bute<br />

directly to <strong>ri</strong>pening by sending photosynthates to the grains. <strong>and</strong> the number <strong>of</strong><br />

nodes at the base <strong>of</strong> the stem from where tillers come out is also almost constant<br />

with a given cultural condition. Therefore, the number <strong>of</strong> nodes. from which<br />

neither tillers nor leaves cont<strong>ri</strong>bute directly" to <strong>ri</strong>pening, is greater in low than<br />

in high-latitude areas. The leaves from such nodes develop du<strong>ri</strong>ng the vegetativelag<br />

phase <strong>and</strong> generally die early. Thus. there are more dead leaves <strong>and</strong> also a<br />

larger number <strong>of</strong> excessively elongated intemodes at the base <strong>of</strong> the stem.<br />

which results in a tall plant <strong>and</strong> a loxver grain-straw ratio in lo\v- than in highlatitude<br />

areas.<br />

This situation is more apparent in the tropics, because long-duration. vegetatively<br />

vigorous va<strong>ri</strong>eties are more popular among farmers. Generally, the total<br />

dryi-matter production is greater but the grain yield is not higher due to a low<br />

grain-straw ratio.<br />

Such a tendency was apparent in Japan 3O years ago (Ishizuka <strong>and</strong> Tanaka,<br />

I956). In northern Japan. such as in Hokkaido, the number <strong>of</strong> leaves on the main<br />

stern <strong>of</strong> medium-duration va<strong>ri</strong>eties was l2, the growth pattern was Type B. <strong>and</strong><br />

the grain yield was not high clue to a relatively’ small total dry-matter production,<br />

although the grain-stratv ratio was reasonable. Due to such a growth pattern, an

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