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Vergara - 1976 - Physiological and morphological adaptability of ri

Vergara - 1976 - Physiological and morphological adaptability of ri

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4 CLINIATE mo) RICE<br />

ag<strong>ri</strong>cultural <strong>and</strong> related sciences. as much as on his own increased expe<strong>ri</strong>ence <strong>of</strong><br />

techniques <strong>and</strong> management.<br />

Of all the factors which affect ag<strong>ri</strong>cultural productivity. the most powerful.<br />

<strong>and</strong> at thc same time least modifiable. is the climate. or its p<strong>ri</strong>ncipal components.<br />

temperature <strong>and</strong> rainfall. Rainfall can. <strong>of</strong> course. be supplemented or replaced<br />

by ir<strong>ri</strong>gation where suitable ivater is available. but even the <strong>ri</strong>ce crop. the traditional<br />

ccreal <strong>of</strong> ir<strong>ri</strong>gation ag<strong>ri</strong>culture. is grown on large areas under rainfcd conditions.<br />

Temperature control is a great deal more costly‘. hence rest<strong>ri</strong>cted to the<br />

cultivation <strong>of</strong> high-cost horticultural products.<br />

On a world scale. temperature presents the most general, <strong>and</strong> the least tractable,<br />

rest<strong>ri</strong>ction on productivity’. although this is less marked in the tropics.<br />

ln general terms. productivity‘ is correlated xivith temperature; subject to the<br />

availability <strong>of</strong> an adequate moisture supply. Most climates are punctuated by<br />

temperature peaks <strong>and</strong> troughs. Their incidence <strong>and</strong> duration. <strong>and</strong> the relationship<br />

to moisture supply. condition the productivity <strong>of</strong> a region. Temperature<br />

in tinze is thus the predominant environmental parameter determining growth".<br />

<strong>and</strong> where thc moisture supply" is irregular. as in a monsoon climate. growing<br />

seasons may be further attenuated.<br />

Cropping seasons, since time imrnemonal, have evolved to fit into the prevailing<br />

climate pattern. To raise the efficiency <strong>of</strong> the system means to utilize to the<br />

maximum the available time, <strong>and</strong>, within its limits, the investment in fertilizers,<br />

cultivation, improved seeds, ir<strong>ri</strong>gation. weed control. <strong>and</strong> plant protection. all<br />

<strong>of</strong> which are components <strong>of</strong> the essential intensification <strong>of</strong> crop production.<br />

Thus temperature, <strong>and</strong> in some important instances temperature in interaction<br />

with the length <strong>of</strong> daylight, is the main controlling factor which determines<br />

whether any or all <strong>of</strong> these investments will be pr<strong>of</strong>itable. Hence the desire to<br />

underst<strong>and</strong> in precise <strong>and</strong> predictable terms the effects <strong>of</strong> temperature <strong>and</strong> day<br />

length on the development <strong>of</strong> crop plants. on their fertilizer <strong>and</strong> water use. on<br />

the tvay they resist or succumb to diseases <strong>and</strong> pests, <strong>and</strong> form. shed, or retain<br />

leaves <strong>and</strong> seeds. Clearly, the (iverall aim must he to devise systems uthich ivill<br />

use to the full the blessings <strong>of</strong> sun energy, but escape its ravages. As ag<strong>ri</strong>culture<br />

advances from p<strong>ri</strong>mitive to intensive levels <strong>of</strong> production. this increasingly<br />

dem<strong>and</strong>s precise adjustment 0f heredity <strong>and</strong> environment, <strong>and</strong> hence a detaiieti<br />

zmderstunciirzg <strong>of</strong>t/re developmental pilrases <strong>of</strong> the plant ‘which determine its productivity‘.<br />

as <strong>of</strong> those phases which are susceptible to injury or breakdown.<br />

It has been claimed that controlled environment facilities could greatly<br />

accelerate this process <strong>of</strong> adjustment. <strong>and</strong> this is very likely indeed. But it must<br />

be understood that the road to “applied phytotrtmics." i.e. to the use <strong>of</strong> a phytotron<br />

in plant selection or agronomy, leads as a rule through a fuller underst<strong>and</strong>ing<br />

<strong>of</strong> the physiologv <strong>of</strong> the plant, i.e. <strong>of</strong> those phases which are c<strong>ri</strong>tical for<br />

development <strong>and</strong> reproduction. The temperatures du<strong>ri</strong>ng both day <strong>and</strong> night,<br />

the daily pe<strong>ri</strong>od <strong>of</strong> illumination, <strong>and</strong> the intensity <strong>and</strong> spectral composition <strong>of</strong><br />

daylight. all have pr<strong>of</strong>ound effects on the growth <strong>of</strong> plants. In the field these

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