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Vergara - 1976 - Physiological and morphological adaptability of ri

Vergara - 1976 - Physiological and morphological adaptability of ri

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PHYSIOLOGICAL AND NIORPIIOLOGICAL ADAPTABILITH’ 69<br />

The plant characters necessary for wide <strong>adaptability</strong> <strong>of</strong> the <strong>ri</strong>ce plant as cited<br />

here deal mainly v<strong>ri</strong>th ir<strong>ri</strong>gated lowl<strong>and</strong> <strong>ri</strong>ce where water is usually not a limiting<br />

factor. The list does not include the other hydro-em<strong>ri</strong>ronmental conditions under<br />

WhlCll the <strong>ri</strong>cc plants are also grown, such as floating-<strong>ri</strong>cc areas (rnorc than I50<br />

cm deep), deep-ivater <strong>ri</strong>ce areas (SO-ISO cm deep), <strong>and</strong> "upl<strong>and</strong>" <strong>ri</strong>ce areas. In<br />

these places. specific <strong>adaptability</strong> is needed.<br />

Drought resistance<br />

Resistance to drought or ability to recover from moisture stress is an important<br />

plant character for specific <strong>adaptability</strong> to rainfed lowl<strong>and</strong> or upl<strong>and</strong> <strong>ri</strong>ce areas.<br />

Different types <strong>of</strong> specific <strong>adaptability</strong> are needed by the <strong>ri</strong>ce plant if it has to<br />

survive in the special habitat where it is being grown, such as under coconut<br />

trees or 5 meters <strong>of</strong> water,<br />

An adaptive mechanism necessary for the floating-<strong>ri</strong>ce va<strong>ri</strong>eties is the ability<br />

to elongate. While insensitivity to photope<strong>ri</strong>tid is necessary‘ under loo/l<strong>and</strong> <strong>and</strong><br />

upl<strong>and</strong> conditions for wide <strong>adaptability</strong>, sensitivity to photopc<strong>ri</strong>od, on the other<br />

h<strong>and</strong>. is required for specific <strong>adaptability</strong> in the floating-<strong>ri</strong>ce areas. The long<br />

growth duration (180 to 200 days) necessary for the floating-<strong>ri</strong>ce areas requires<br />

sensitivity to photopc<strong>ri</strong>od. since there are no known photopc<strong>ri</strong>od-insensitive<br />

va<strong>ri</strong>eties with groivth duration longer than 150 day's.<br />

The va<strong>ri</strong>eties capable <strong>of</strong> producing very high grain yields under conditions <strong>of</strong><br />

widely: fluctuating seasonal environments have many differences in growth form<br />

<strong>and</strong> phyisitiltigy. Although one can define certain characte<strong>ri</strong>stics <strong>of</strong> plants<br />

adapted to specific environments, it is difficult if not impossible to define all the<br />

possible combinations <strong>of</strong> characte<strong>ri</strong>stics necessary to provide good general<br />

<strong>adaptability</strong> to a utidely fluctuating set <strong>of</strong> seasons (Finlay <strong>and</strong> Wilkinson, 1963).<br />

Hence. the climatic factors have been divided into day length, wind, solar radiation,<br />

precipitation, <strong>and</strong> temperature; <strong>and</strong> no attempt is made in this paper to<br />

define all the possible combinations to provide general <strong>adaptability</strong>‘.<br />

Most <strong>of</strong> the discussion will involve specific rather than general <strong>adaptability</strong>.<br />

DAY LENG TI I<br />

Like the wild <strong>ri</strong>ce, the early cultivated-<strong>ri</strong>ce va<strong>ri</strong>eties were grown in lovv-ly-‘ing<br />

areas where water is available; <strong>and</strong> in many cases it is the only crop that can be<br />

grown. because <strong>of</strong> too much water To avoid the cumbersome method <strong>of</strong> harvesting<br />

<strong>ri</strong>ce while the water is still deep or the weather is too wet for drying <strong>of</strong> grain,<br />

va<strong>ri</strong>eties have been selected that mature when the water has receded or when the<br />

soil is starting to dry out.<br />

In photope<strong>ri</strong>od-sensitivc va<strong>ri</strong>eties. delay in transplanting. resulting from late<br />

rains, will not affect grain yields under lowl<strong>and</strong> conditions. In this sense the<br />

photopc<strong>ri</strong>odscnsitive or the long growtlrduration va<strong>ri</strong>eties have a more flexible<br />

seedling stage.<br />

Before the introduction <strong>of</strong> the high-yielding <strong>ri</strong>cc va<strong>ri</strong>eties in the tropics. the

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