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Revision of Passiflora Subgenus Decaloba ... - Passion Flowers

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283<br />

shallowly trilobed leaves (commonly less than half the distance from the leaf<br />

outline to the leaf base) with the length <strong>of</strong> the central lobe <strong>of</strong>ten greatly exceeding<br />

that <strong>of</strong> the lateral lobes and an angle between the lateral veins that is frequently<br />

between 40 and 80°. The lateral lobes are also oblong to elliptic. The broadly<br />

capitate petiolar nectaries are commonly positioned on the distal half <strong>of</strong> the<br />

petiole, <strong>of</strong>ten over 0.60 the distance from the base to the apex <strong>of</strong> the petiole.<br />

The leaf bases are <strong>of</strong>ten cuneate to acute but rarely cordate. In México and<br />

Central America, <strong>Passiflora</strong> suberosa ssp. litoralis is <strong>of</strong>ten found in high elevation<br />

(1000 -3000 m) moist pine and oak forests along streams and rivers, but it may<br />

also occur in very dry forests with cacti (e.g., Cephalocereus forests <strong>of</strong><br />

Tehuacán) and other species common in matorral vegetation (e.g., Tamaulipan<br />

matorral).<br />

On the western side <strong>of</strong> South America (Colombia to Perú and Argentina), P.<br />

suberosa ssp. litoralis possesses leaves very much like those <strong>of</strong> the<br />

México/Central American variant, but the petiolar nectaries may be more discoid<br />

and are commonly positioned on the proximal half <strong>of</strong> the petiole. The lateral<br />

lobes are commonly distinctly ovate and diverge at an angle <strong>of</strong> 80-100°. The leaf<br />

bases are also distinctly cordate. In this region it is found in low (near sea level<br />

and on cliffs above the sea) to high (to 3,000 m) elevation moist forests<br />

commonly along streams and rivers, but it also occurs in tropical dry forests.<br />

On the eastern side <strong>of</strong> South America, in southeastern Brazil, the leaves are<br />

commonly trilobed but may also have unlobed, bilobed or trilobed leaves present<br />

on the same plant. The petiolar nectaries are commonly discoid and positioned

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