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Operational Plan for the Restoration of Diadromous Fishes to the ...

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4) The model developed by Solow and Costello (2004) has a Poisson<br />

distribution with an annual introduction rate described by μ t = exp( β 0 + β 1t<br />

),<br />

where β 0 and β1 were estimated through a maximum-likelihood method and<br />

are reported above. The Poisson distribution is useful in describing counts<br />

from rare events over a continuous timeframe. A Poisson probability density<br />

function (PDF) can be developed from <strong>the</strong> annual introduction rate that<br />

describes <strong>the</strong> probability <strong>of</strong> <strong>the</strong> number <strong>of</strong> discoveries one could expect <strong>for</strong><br />

that year. The PDF can be used <strong>to</strong> estimate <strong>the</strong> probability <strong>of</strong> <strong>the</strong> number <strong>of</strong><br />

discoveries occurring in any one year by:<br />

x<br />

λ<br />

P( x)<br />

= x<br />

e x!<br />

where x is <strong>the</strong> number <strong>of</strong> discoveries in an interval and λ is <strong>the</strong> mean rate<br />

over <strong>the</strong> interval. For this application, <strong>the</strong> mean introduction rate in 2007 <strong>of</strong><br />

2.64 introductions was used <strong>to</strong> determine <strong>the</strong> current probability <strong>for</strong> humancaused<br />

introductions. The probability <strong>of</strong> nor<strong>the</strong>rn pike dispersal in any one<br />

year was constructed by taking <strong>the</strong> difference (subtracting <strong>the</strong> two mean rates<br />

<strong>for</strong> 2007) <strong>of</strong> <strong>the</strong> combined and introduced models. The probability functions<br />

<strong>for</strong> human-caused introduction (smooth line) and natural dispersal (dashed<br />

line) are plotted in Figure 12.<br />

Annual Probability<br />

0.4000<br />

0.3500<br />

0.3000<br />

0.2500<br />

0.2000<br />

0.1500<br />

0.1000<br />

0.0500<br />

0.0000<br />

0 1 2 3 4 5 6 7 8 9 10 11<br />

Number <strong>of</strong> Introductions<br />

Human-caused Dispersal<br />

Figure 12 Probability density functions <strong>for</strong> human-caused introductions and dispersal <strong>of</strong> nor<strong>the</strong>rn pike<br />

Each curve in Figure 12 represents <strong>the</strong> probability <strong>of</strong> an introduction occurring<br />

in any one year in Maine. Inspection <strong>of</strong> <strong>the</strong> PDF <strong>for</strong> each case provides some<br />

insights on <strong>the</strong> risk <strong>of</strong> each pathway:<br />

a. The probability <strong>of</strong> having no introductions (maintaining <strong>the</strong> current<br />

condition) is more than four times more likely under natural dispersal ( ρ =<br />

0.32) than under human-caused introductions ( ρ =0.07). Natural dispersal<br />

PRFP Page 249

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