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Control of Virus Diseases of Plants 151challenge (Verma and Mukerji, 1975; Verma and Awasthi, 1979, 1980; Vermaet al., 1984; Verma et al., 1995a; Prasad et al., 1995; Verma et al., 1996). Such<strong>plant</strong> <strong>products</strong> have been called systemic resistance <strong>in</strong>ducers (SRIs).2. Basic prote<strong>in</strong>s from <strong>plant</strong>s that function by <strong>in</strong>activat<strong>in</strong>g ribosomes of thehost and have been called RIPs (Barbieri et al., 1993, 2003; Van Damme et al.,2001).7.2 Systemic Resistance Inducers (SRIs)Induction of systemic resistance by <strong>plant</strong> extracts has been reviewed fromtime to time (Verma, 1985; Verma and Prasad, 1992; Verma et al., 1995a; Vermaet al., 1998). Physico-chemical characteristics of systemic resistance <strong>in</strong>ducersfrom <strong>plant</strong>s such as Boerhaavia diffusa (Verma and Awasthi, 1979, 1980; Srivastava,1995); Mirabilis jalapa (Verma and Kumar, 1980); Cuscuta reflexa(Awasthi, 1981); Clerodendrum aculeatum (Verma et al., 1984; Verma et al., 1996;Kumar et al., 1997; Srivastava et al., 2008); Bouga<strong>in</strong>villea spectabilis (Vermaet al., 1985; Verma and Dwivedi, 1984; Srivastava, 1995); Pseuderanthemumatropurpureum (Verma et al., 1985) have been studied <strong>in</strong> some detail (Table7.1).Induced resistance operates through the activation of <strong>natural</strong> defencemechanisms of the host <strong>plant</strong>. Extracts from br<strong>in</strong>jal (Verma and Mukherjee,1975) and a few other higher <strong>plant</strong>s such as Boerhaavia diffusa (Verma et al.,1979), Bouga<strong>in</strong>villea (Verma and Dwivedi, 1984), Clerodendrum (Vermaet al., 1996; Kumar et al., 1997) and Datura (Verma et al., 1982) <strong>in</strong>duce systemicresistance to viral multiplication <strong>in</strong> <strong>plant</strong>s. The active <strong>products</strong> present <strong>in</strong> theseTable 7.1. Characteristics of systemic resistance <strong>in</strong>ducers obta<strong>in</strong>ed from some higher <strong>plant</strong>s.CharacteristicsSRI-yield<strong>in</strong>g <strong>plant</strong>sBD CA/CI BS PA CR MJSource Root Leaf Leaf Leaf Leaf LeafThermal <strong>in</strong>activation 80 90 80 0 70 90po<strong>in</strong>ts (°C )Nature Glyco-prote<strong>in</strong> Basic Prote<strong>in</strong> Prote<strong>in</strong> 0 Prote<strong>in</strong> Prote<strong>in</strong>Molecular weight 30 34/29 and 34 28 0 14–18 24(kDa)Active aga<strong>in</strong>st TMV TMV TMV TMV TMV TMVSHRV SHRV SHRV SHRV SHRV SHRVGMV GMV CGMMV CGMMV GMV PVYPLRV TmYMVBD = Boerhaavia diffusa; CA = Clerodendrum aculeatum; CI = C. <strong>in</strong>erme; BS = Bouga<strong>in</strong>villea spectabilis;PA = Pseuderanthemum atropurpureum; CR = Cuscuta refl exa; MJ = Mirabilis jalapa. TMV = Tobaccomosaic virus; SHRV = Sunnhemp rosette virus; GMV = Gomphrena mosaic virus; TRSV = TobaccoR<strong>in</strong>gspot virus; TmYMV = Tomato Yellow Mosaic virus; CGMMV = Cucumber green mottle mosaic virus;PVY = Potato virus Y; PLRV = Papaya leaf reduction virus.

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