The Descent of Man

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ledge that the blood of some more humble creature flows in his veins. For my own part I would as soon be descended from that heroic little monkey, who braved his dreaded enemy in order to save the life of his keeper, or from that old baboon, who descending from the mountains, carried away in triumph his young comrade from a crowd of astonished dogs—as from a savage who delights to torture his enemies, offers up bloody sacrifices, practices infanticide without remorse, treats his wives like slaves, knows no decency, and is haunted by the grossest superstitions. Man may be excused for feeling some pride at having risen, though not through his own exertions, to the very summit of the organic scale; and the fact of his having thus risen, instead of having been aboriginally placed there, may give him hope for a still higher destiny in the distant future. But we are not here concerned with hopes or fears, only with the truth as far
as our reason permits us to discover it; and I have given the evidence to the best of my ability. We must, however, acknowledge, as it seems to me, that man with all his noble qualities, with sympathy which feels for the most debased, with benevolence which extends not only to other men but to the humblest living creature, with his god-like intellect which has penetrated into the movements and constitution of the solar system—with all these exalted powers—Man still bears in his bodily frame the indelible stamp of his lowly origin. SUPPLEMENTAL NOTE. ON SEXUAL SELECTION IN RELATION TO MONKEYS. Reprinted from NATURE, November 2, 1876, p. 18.
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Work reproduced with no editorial r
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INTRODUCTION. The nature of the fol
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form, we are deprived of the weight
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This work contains hardly any origi
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probably never have completed it. A
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aces of man. But owing to the lengt
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must be classed as doubtful species
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ain of man has its analogy in that
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chemical reagent.) of their tissues
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prove how similar the nerves of tas
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idem in Cynocephalo. Illustrissimus
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First visceral arch. g. Second visc
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oth drawings removed. I was directe
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will conclude with a quotation from
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are useless, or nearly useless, and
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vol. ii pp. 317 and 397. See also '
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Some few persons have the power of
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attention towards them, could recov
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large and heavy birds, which, from
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and Mr. E. Ray Lankester informs me
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een sent me, the projection is so l
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e rapidly drawn across the whole ey
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white races. See his paper, 'Medico
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the body are the rudiments of the u
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naked, like the inferior surfaces o
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thropological Review, July 1867, p.
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pendage is a rudiment, we may infer
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Petersbourg,' tom. xii. 1867, p. 44
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pont found thirty per cent. of perf
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y Prof. Turner, has been expressly
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inheritance, that is, on parts acqu
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etc., is utterly inexplicable. (56.
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seal, bat, reptile, etc., can at fi
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Variability of body and mind in man
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much diversity in the teeth as in t
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The variability or diversity of the
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wonderfully complex combination of
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has never long been controlled, eit
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If then in outward manner, form, an
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continued use or disuse of parts. T
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Military and Anthrop. Statistics,'
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arrive at any precise result. Dr. B
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their coats. When one kidney ceases
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is hereditary; there is really some
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Variation of Animals under Domestic
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phere, they have acquired chests an
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served that they still inherited th
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idiots, as described in Vogt's memo
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Many of the cases to be here given,
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mammae (of course rudimentary) in t
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mestication why I now place little
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into the body of the uterus." The a
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Professor Canestrini, after discuss
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pecially, as Canestrini has observe
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nine character, as Owen (42. 'Anato
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to an ape-like progenitor. He who r
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Dr. Murie and Mr. St. George Mivart
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ing muscles in the lower animals. (
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muscles of the human neck, shoulder
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CORRELATED VARIATION. In man, as in
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RATE OF INCREASE. Civilised populat
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suckling their infants during a lon
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support a wife; and they generally
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pan) it has been intentionally enco
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elephant, the slowest breeder of al
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ther occasionally or habitually, ha
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smallest proficiency in it might re
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only have endowed the savage with a
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with precision, or to form a flint
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and, as I have myself seen, are qui
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sume his erect attitude, which form
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progression of the higher apes, to
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in structure more nearly to the bip
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teeth; but as they gradually acquir
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tles. (78. Dujardin, 'Annales des S
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inhabitants of Britain have "much m
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many facts shew how easily the skul
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was 3.15 and the latter 4.3 inches
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ates,' vol. iii. p. 619.) than thos
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have led to the denudation of his b
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pologie,' 1872; 'La Constitution de
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unneus is formed of eleven vertebra
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epilepsy in guinea-pigs, and likewi
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gether. Independently of correlatio
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secondly, that natural selection ha
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the injurious will be thus eliminat
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Judging from the habits of savages
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adduces the naked and unprotected s
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from his fellow-men. No country in
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descent from some lower form; but i
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Differences of this kind between th
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somewhat fewer instincts than those
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Although the first dawnings of inte
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a reflex action, yet it is not impr
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his work by practice; a beaver, on
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dotes have been published on the lo
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maternal affection exhibited in the
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in the old baboon. Monkeys will als
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We will now turn to the more intell
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frightened, they soon approached, h
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ents, and sometimes of other birds;
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easily approached. Mr. Bartlett has
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The IMAGINATION is one of the highe
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liberate, and resolve. It is a sign
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fishes, that he was sometimes compl
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I have seen, as I daresay have othe
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ly as would a philosopher in his lo
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The following cases relate to dogs.
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writers even yet deny that the high
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such aphorisms, but they are almost
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have come into play. I have receive
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Mus coninga to its superior cunning
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were employed as implements; but th
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implement for a special purpose is
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It would be very difficult for any
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never reflects on his past pleasure
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hypothesis is a false one." (46. Th
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movements of the muscles of the fac
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ii. p. 309. Dr. A. Moschkau informs
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olog., tom. x. p. 119.) has proved,
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cal cadences, that is in singing, a
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early progenitor of man must have b
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partly on the disposition of the mi
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certain sounds with certain general
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larly constructed, these being used
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often remain as the rudiments of an
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the Lapponian, and many of the Amer
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SENSE OF BEAUTY. This sense has bee
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irregular intervals are highly disa
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most remarkable and typical differe
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As Mr. M'Lennan (75. 'The Worship o
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dowed with some form of life, and w
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eligious rites; and Jemmy Button, w
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lood-loving god; the trial of innoc
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sense or conscience is by far the m
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man is a social animal ('Psychologi
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these feelings and services are by
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would acquire exactly the same mora
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tain stronger or more enduring inst
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first discovers an enemy, warns the
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drive away a third stallion from a
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they actually sympathise in the pai
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his companions. Mr. Blyth, as he in
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to escape, after, as the surgeon th
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teach him silence and obedience. Mr
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than a thousand miles, could have f
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duals which took the greatest pleas
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Bain's Mental and Moral Science,' 1
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It is, however, impossible to decid
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east against the wires of her cage,
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of the year during the whole day, m
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same manner as with the lower anima
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judgment of his fellow-men, and unf
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y the same instinctive motive, whic
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their comrades; and surely their co
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actions, yet it is untenable, that
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felt except in the presence of dang
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pressions of past hunger, vengeance
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will surely come. He will then feel
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een stirred to the bottom of his so
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hesitation." (29. E.B. Tylor, in 'C
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he has no such sympathy, and if his
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tant. But they are practised almost
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that an Indian Thug conscientiously
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to your enemy has rarely been thoug
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unmarried females. How slowly it sp
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I have entered into the above detai
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the two do not easily co-exist in t
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and to take as the standard of mora
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influence in the burning sense of s
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supported as they are by reason, sh
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spreads through instruction and exa
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are likewise transmitted. That the
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some feeling of sympathy, and have
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to the feelings caused by other pow
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sounds had never crossed their mind
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Marcus Aurelius,' etc., p. 139.)—
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difications through natural selecti
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er of offspring. The tribes, which
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tedly it would be interesting to tr
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ed was found to be valuable, the de
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course of time it would, judging fr
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nevolent actions. Habits, moreover,
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We may therefore conclude that prim
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morality will certainly give an imm
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Fuego, it struck me that the posses
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a vigorous state of health. We civi
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In every country in which a large s
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doubt wealth when very great tends
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ges. (13. Quatrefages, 'Revue des C
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higher degree by their works than b
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men rarely marry; both suffer from
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in comfort. Those who marry early p
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towns than in rural districts, "and
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Report of Births, Deaths, etc., in
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moral qualities. (25. Dr. Duncan re
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mind and body. But development of a
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them. In Spain alone some of the be
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Natural selection follows from the
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causes of progress seem to consist
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Botocudos, who inhabit the finest p
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Testament.") and not admit that alm
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eady become somewhat advanced. Look
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Even if it be granted that the diff
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kingdom. (1. Isidore Geoffroy St.-H
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slaves. They move the eggs of their
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vol. ii. 1840, p. 87.) Classificati
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We can further see why a great amou
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apparently follow from their differ
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as well as by various drugs, and ot
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Richtung der Haare,' etc., Muller's
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It would, however, be rash to trust
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Although, as we have now seen, man
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tion of the Simiadae. This family i
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the brain, does not admit the exist
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and Platyrrhine monkeys, with their
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1868, s. 61. Also his 'Naturliche S
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We are far from knowing how long ag
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hous apes, as Professor Schaaffhaus
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Most of the remnants survive on isl
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eason to believe, that no true bird
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evidently a widely-diffused remnant
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the satisfaction of seeing, in Apri
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then be justified in believing that
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period the uterus was double; the e
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counter a singular difficulty. In t
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The possession by male mammals of f
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and in the case of the Marsupials,
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sionally to become so well develope
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great kingdom of the Vertebrata it
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in two weeks; those of the fowl in
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quality. The world, it has often be
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alists are practically guided by th
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etween the races, we must make some
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tions of the brain. (3. See, for in
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tian caves of Abou-Simbel, M. Pouch
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low temperature, or any considerabl
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is, whether any of them are descend
Page 477 and 478:
e whether the races of men, when cr
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and mulattoes, have a maxim that mu
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laws, for instance, that of the une
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e produced at the rarest intervals.
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completely black or completely whit
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dians, and Europeans; and judging f
Page 489 and 490:
But the most weighty of all the arg
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those forms which at present appear
Page 493 and 494:
The question whether mankind consis
Page 495 and 496:
her all the domestic races of the d
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ner with the improved breeds of cat
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plainly retained, in certain bones
Page 501 and 502:
gesture-language, by the same expre
Page 503 and 504:
The same argument may be applied wi
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in any extreme degree to those poss
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With our domestic animals a new rac
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speak a word of the language of a l
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down the numbers of each savage tri
Page 513 and 514:
extinction, believed that changed h
Page 515 and 516:
who were in 1832 transported to Fli
Page 517 and 518:
Similar facts have been observed in
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unproductiveness of the women, and
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ious places, in order to be educate
Page 523 and 524:
ding to Dr. Ruschenberger of the U.
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the natives, and they soon became "
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tants of the Valley of Nepal, which
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aborigines of islands have suffered
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many years together in their own co
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We can see why it is that aborigine
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y the House of Commons, May 29, 186
Page 537 and 538:
certain the precise causes and thei
Page 539 and 540:
But as with our domesticated animal
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posed to be more influential in mod
Page 543 and 544:
negro seems to be partly inherent,
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improbable. Consequently I endeavou
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late Dr. Daniell, who had long live
Page 549 and 550:
ed by a medical man, that some year
Page 551 and 552:
cording to the position of the vall
Page 553 and 554:
the Fuegians on board the "Beagle"
Page 555 and 556:
e of the Mandans, about one in ten
Page 557 and 558:
morphic, which have remained extrem
Page 559 and 560:
NOTE ON THE RESEMBLANCES AND DIFFER
Page 561 and 562:
B. x. 1868.) on the cerebral convol
Page 563 and 564:
ule, in man, the temporo-occipital
Page 565 and 566:
paper on this subject the latter wr
Page 567 and 568:
1862.); and more or less obliterate
Page 569 and 570:
In the paper to which I have referr
Page 571 and 572:
This statement was a strictly accur
Page 573 and 574:
mitting the similarity between the
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giques de Gratiolet' ('Mem. de la S
Page 577 and 578:
made the subject of renewed investi
Page 579 and 580:
pal sulci of the frontal, parietal,
Page 581 and 582:
with no sulci but the sylvian and t
Page 583 and 584:
excellent author of one of the most
Page 585 and 586:
several forms of inheritance—Caus
Page 587 and 588:
here concern us. There are, however
Page 589 and 590:
are generally confined to the lower
Page 591 and 592:
find the female, he requires organs
Page 593 and 594:
locomotive organs more highly devel
Page 595 and 596:
vocal or instrumental music—and t
Page 597 and 598:
state of nature, have by a long sel
Page 599 and 600:
Mr. Swaysland of Brighton has been
Page 601 and 602:
young—a period which must be dete
Page 603 and 604:
former, there can be little doubt,
Page 605 and 606:
so simple and short an affair as mi
Page 607 and 608:
NUMERICAL PROPORTION OF THE TWO SEX
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ment of doubt; for it is a well-asc
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to lead to effective sexual selecti
Page 613 and 614:
We will first briefly run through t
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is polygamous, but, except in his g
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far as I can learn, the sexes rarel
Page 619 and 620:
differences, are polygamous, but Mr
Page 621 and 622:
irds, whether the male tragopan (on
Page 623 and 624:
females, which must pair with the l
Page 625 and 626:
apparently of Batrachians. Througho
Page 627 and 628:
paratively passive, generally exert
Page 629 and 630:
should retain the same habit; and t
Page 631 and 632:
dings of the Royal Society,' vol. x
Page 633 and 634:
very many instances widely differen
Page 635 and 636:
tion. Nevertheless, they may become
Page 637 and 638:
males, and the latter the more attr
Page 639 and 640:
al selection for the sake of protec
Page 641 and 642:
certain stags—has been carried to
Page 643 and 644:
hackles, comb, spurs, voice, and ev
Page 645 and 646:
developed atoms, which are transmit
Page 647 and 648:
feather is transversely marked by n
Page 649 and 650:
change the colour of the coat durin
Page 651 and 652:
in the English Carrier pigeon, and
Page 653 and 654:
There is one difficult question whi
Page 655 and 656:
markable how very closely he sugges
Page 657 and 658:
fer much in constitution before the
Page 659 and 660:
acters of the adults must have been
Page 661 and 662:
the roebuck, to ten, twelve or even
Page 663 and 664:
horns of the female: see also his p
Page 665 and 666:
veloped later in life in this breed
Page 667 and 668:
in the common peacock; but M. Hegt
Page 669 and 670:
which the characters follow our two
Page 671 and 672:
pear late in life in one sex are tr
Page 673 and 674:
the class previously alluded to, in
Page 675 and 676:
laced plumage of the Sebright banta
Page 677 and 678:
the reindeer, in which both sexes b
Page 679 and 680:
either or both sexes have been affe
Page 681 and 682:
y acquiring structures, such as gre
Page 683 and 684:
have not frequently been accumulate
Page 685 and 686:
As no one, as far as I can discover
Page 687 and 688:
the female births have exceeded the
Page 689 and 690:
ce to show that, at almost every st
Page 691 and 692:
It seems at first sight a mysteriou
Page 693 and 694:
important though not the sole facto
Page 695 and 696:
failed to produce offspring. During
Page 697 and 698:
lieve that females are produced in
Page 699 and 700:
as well as some of the following re
Page 701 and 702:
females: the natives believe that t
Page 703 and 704:
Sweden,' 1867, pp. 12, 132.), that
Page 705 and 706:
one. It should, however, be borne i
Page 707 and 708:
Infertile females might readily be
Page 709 and 710:
than flight." He then adds, that by
Page 711 and 712:
nion appears to be that the sexes a
Page 713 and 714:
in number; but three South African
Page 715 and 716:
ed from the eggs and caterpillars a
Page 717 and 718:
maturity, when the sexes are ready
Page 719 and 720:
or varieties of moths (Heterocera)
Page 721 and 722:
number, whatever the proportions ma
Page 723 and 724:
les, and "two or three are often fo
Page 725 and 726:
exceeded the females in number; in
Page 727 and 728:
and thus we see how ignorant we are
Page 729 and 730:
males; but he suggests that this pr
Page 731 and 732:
nious manner. "Let us for the purpo
Page 733 and 734:
monstratively fixed, it is quite cl
Page 735 and 736:
servations apply to the whole archi
Page 737 and 738:
and kept from the use of spirits. H
Page 739 and 740:
From the several foregoing cases we
Page 741 and 742:
oons, it is conceivable that a male
Page 743 and 744:
acquired before maturity—Spiders,
Page 745 and 746:
condition of others, and the low me
Page 747 and 748:
devourers that they are distasteful
Page 749 and 750:
the Molluscoida of some authors), f
Page 751 and 752:
strong and healthy individual disap
Page 753 and 754:
influential to a certain extent; fo
Page 755 and 756:
organised creatures this is extreme
Page 757 and 758:
ting a partner, or for the individu
Page 759 and 760:
the number of the smelling-threads,
Page 761 and 762:
chelae of the male. N.B.—The arti
Page 763 and 764:
cording to a statement quoted by Mi
Page 765 and 766:
certainly suffice to hold the femal
Page 767 and 768:
fe. This fact shews that in the Amp
Page 769 and 770:
sexes of the higher animals so ofte
Page 771 and 772:
the larger size of their chelae. In
Page 773 and 774:
ordinary coloration of the genus to
Page 775 and 776:
each other in the size and length o
Page 777 and 778:
and too small for her to catch with
Page 779 and 780:
legs belonging either to one of the
Page 781 and 782:
213.) The ocelli are absent in the
Page 783 and 784:
hed, so that he is thus enabled to
Page 785 and 786:
the male of Crabro cribrarius (Fig.
Page 787 and 788:
ation (10. E. Doubleday, 'Annals an
Page 789 and 790:
mals, we can understand why she has
Page 791 and 792:
create a large number of offspring
Page 793 and 794:
ger than the females. The explanati
Page 795 and 796:
quite black, and the females obscur
Page 797 and 798:
33) that, "the captured flies utter
Page 799 and 800:
ORDER: HOMOPTERA. Every one who has
Page 801 and 802:
temdecim,' by Dr. Hartman.), "the d
Page 803 and 804:
unmusical even to the human ear, so
Page 805 and 806:
d'Anat. Comp.' (French translat.),
Page 807 and 808:
Coleoptera. But Dr. Gruber further
Page 809 and 810:
wing-cover, which underlies the oth
Page 811 and 812:
In the last and third family, namel
Page 813 and 814:
Cape of Good Hope these insects mak
Page 815 and 816:
yed by the Homoptera. (43. Landois
Page 817 and 818:
p. 427; for crickets, p. 445.) With
Page 819 and 820:
with splendid green, blue, yellow,
Page 821 and 822:
sub-families, the males on first em
Page 823 and 824:
mes two chasing one female, and con
Page 825 and 826:
In this Order slight differences in
Page 827 and 828:
for the possession of the females;
Page 829 and 830:
worker-bees is a much more difficul
Page 831 and 832:
some degree probable in those group
Page 833 and 834:
one species both are of a beautiful
Page 835 and 836:
genus Onthophagus, there are specie
Page 837 and 838:
espect amongst well-developed indiv
Page 839 and 840:
projection in the same situation. S
Page 841 and 842:
species of Bledius (Fig. 23), also
Page 843 and 844:
187.) enclosed two males with one f
Page 845 and 846:
provided or degenerate males. Altho
Page 847 and 848:
tance of several feet or even yards
Page 849 and 850:
indebted to Mr. E.W. Janson, for in
Page 851 and 852:
[Fig.26. Hind-leg of Geotrupes ster
Page 853 and 854:
that from the noise thus produced b
Page 855 and 856:
me number of females; in order, the
Page 857 and 858:
ut the chief difference is that the
Page 859 and 860:
devour a large beetle, would be fri
Page 861 and 862:
illiant in the males—Examples—N
Page 863 and 864:
The Ageronia feronia makes a noise
Page 865 and 866:
Every one must have admired the ext
Page 867 and 868:
don,' 1869, p. 278.) In nine of the
Page 869 and 870:
sexes. In a few species, for instan
Page 871 and 872:
the species belongs. Hence in most
Page 873 and 874:
follow exactly the same habits of l
Page 875 and 876:
in which the sexes differ greatly i
Page 877 and 878:
As Mr. Walsh has remarked to me, th
Page 879 and 880:
mies; nevertheless, as the hind-win
Page 881 and 882:
The bright colours of many butterfl
Page 883 and 884:
than the lower. Hence the lower sur
Page 885 and 886:
"holding their wings quite erect ov
Page 887 and 888:
fully explained. The white female o
Page 889 and 890:
We know that ants and certain Lamel
Page 891 and 892:
other band, the females habitually,
Page 893 and 894:
I have as yet only referred to the
Page 895 and 896:
it is the females which "are orname
Page 897 and 898:
least gaudy. The male of Papilio se
Page 899 and 900:
sexes, though arising and perfected
Page 901 and 902:
elieves that the difference between
Page 903 and 904:
they would tend to counteract each
Page 905 and 906:
orange wing-tips of the male; conse
Page 907 and 908:
Soc.' vol. xxiii. 1862, p. 495.), w
Page 909 and 910:
valuable, as Mr. Riley here discuss
Page 911 and 912:
mitted to, and developed in, the ma
Page 913 and 914:
length, transversely banded with bl
Page 915 and 916:
p. lxxx.) it was supported by vario
Page 917 and 918:
SUMMARY AND CONCLUDING RE- MARKS ON
Page 919 and 920:
make the loudest or most continuous
Page 921 and 922:
tected species inhabiting the same
Page 923 and 924:
the more ornamented, and departs mo
Page 925 and 926:
the Coleoptera, however, as before
Page 927 and 928:
ed, in all these respects between b
Page 929 and 930:
upper outer surface of their pector
Page 931 and 932:
her hiding-place and surveys the ne
Page 933 and 934:
and many so injure each other as to
Page 935 and 936:
projection, and the teeth grow into
Page 937 and 938:
tish Fishes,' vol. ii. 1836, p. 416
Page 939 and 940:
cal fishes differ sexually in colou
Page 941 and 942:
ightest, to conceive the admixture
Page 943 and 944:
In a siluroid fish, inhabiting the
Page 945 and 946:
ny of the Chromidae, for instance i
Page 947 and 948:
and iridescent. (24. 'The American
Page 949 and 950:
myriad ova with which they are lade
Page 951 and 952:
To return to our more immediate sub
Page 953 and 954:
animals, splendid colours may be th
Page 955 and 956:
they were thus rendered highly cons
Page 957 and 958:
tain fishes, moreover, can through
Page 959 and 960:
Certain fishes, belonging to severa
Page 961 and 962:
the spawning-season than at any oth
Page 963 and 964:
ather brighter than the females. Th
Page 965 and 966:
the same sex, and accumulated throu
Page 967 and 968:
May 1870, p. 40.) From this stateme
Page 969 and 970:
ted, though lurid tints, and these
Page 971 and 972:
that as soon as he saw its happy se
Page 973 and 974:
to sit in the evening to listen to
Page 975 and 976:
than the females: during the pairin
Page 977 and 978:
snakes of N. America, the male of w
Page 979 and 980:
They are not known to fight togethe
Page 981 and 982:
of the first coral- snake which I s
Page 983 and 984:
that the rattle may not be of use t
Page 985 and 986:
[Fig.33. Sitana minor. Male with th
Page 987 and 988:
likewise have a low crest running a
Page 989 and 990:
Fig. 36. Chamaeleo Owenii. Upper fi
Page 991 and 992:
labris there is a still greater dif
Page 993 and 994:
played almost as important a part w
Page 995 and 996:
during the summer months is confine
Page 997 and 998:
y Mr. J. Jenner Weir that the bird-
Page 999 and 1000:
'Natural History of Ireland: Birds,
Page 1001 and 1002:
ta); and where these fights have oc
Page 1003 and 1004:
namely spurs, which can be used wit
Page 1005 and 1006:
have fought; and the black-cocks "m
Page 1007 and 1008:
has been seen with five spurs. The
Page 1009 and 1010:
s. 740. See also on this bird Azara
Page 1011 and 1012:
ious other birds, they are generall
Page 1013 and 1014:
me more brilliant during the breedi
Page 1015 and 1016:
sing noise made by some nestling-bi
Page 1017 and 1018:
irds during his whole life, asserts
Page 1019 and 1020:
is clear, for a sterile, hybrid can
Page 1021 and 1022:
has also been argued, that the song
Page 1023 and 1024:
'Todd's Cyclopaedia of Anatomy and
Page 1025 and 1026:
In some birds the vocal organs diff
Page 1027 and 1028:
hold water, but is connected with t
Page 1029 and 1030:
is furnished with an additional pai
Page 1031 and 1032:
lack-cock certainly serves as a cal
Page 1033 and 1034:
directly to the place where the mal
Page 1035 and 1036:
Grouse, Richardson, 'Fauna Bor. Ame
Page 1037 and 1038:
webs being strongly bound together.
Page 1039 and 1040:
known to make a humming noise whils
Page 1041 and 1042:
Sclater, in 'Proceedings, Zoologica
Page 1043 and 1044:
the raw meat and reeking liver take
Page 1045 and 1046:
y summons," and when they approach
Page 1047 and 1048:
tain Stokes has described the habit
Page 1049 and 1050:
they often swell and assume vivid t
Page 1051 and 1052:
sexual selection of these ornamenta
Page 1053 and 1054:
hers of this kind occur in the tail
Page 1055 and 1056:
consequently tending to vary in the
Page 1057 and 1058:
ald, "and of a rich cobalt blue, cr
Page 1059 and 1060:
very rare colour in terrestrial spe
Page 1061 and 1062:
ing this period on the head of the
Page 1063 and 1064:
Fourthly, there are birds the sexes
Page 1065 and 1066:
fluence on any other characters; an
Page 1067 and 1068:
(81. On the moulting of the ptarmig
Page 1069 and 1070:
undergo a double moult, keep their
Page 1071 and 1072:
with some proper to the female; and
Page 1073 and 1074:
cal Biography,' vol. i. pp. 174, 22
Page 1075 and 1076:
trains, and strutting about in all
Page 1077 and 1078:
finement in the Malay Archipelago,
Page 1079 and 1080:
attitude the ocelli over the whole
Page 1081 and 1082:
The ocelli on the wing-feathers are
Page 1083 and 1084:
dary feathers, when they are all ex
Page 1085 and 1086:
also, finds that all male birds wit
Page 1087 and 1088:
once than otherwise would be the ca
Page 1089 and 1090:
Gould's 'Handbook to the Birds of A
Page 1091 and 1092:
when we see a peacock strutting abo
Page 1093 and 1094:
'Nat. Hist. Lib.: Birds,' vol. xiv.
Page 1095 and 1096:
CHAPTER XIV. BIRDS—continued. Cho
Page 1097 and 1098:
LENGTH OF COURTSHIP. The lengthened
Page 1099 and 1100:
they may be seen running about "lik
Page 1101 and 1102:
owers of the Bower- birds are the r
Page 1103 and 1104:
suspect, improbable as this will at
Page 1105 and 1106:
it probable that most of them were
Page 1107 and 1108:
mate, and so for several times foll
Page 1109 and 1110:
irds, are always seen during the sp
Page 1111 and 1112:
the authority of the Hon. and Rev.
Page 1113 and 1114:
sion. (9. I am indebted to Prof. Ne
Page 1115 and 1116:
homes after an interval of nine mon
Page 1117 and 1118:
as is proved by the strong and perm
Page 1119 and 1120:
lack- headed. This bullfinch was a
Page 1121 and 1122:
Mr. Gould states that certain hummi
Page 1123 and 1124:
vered;" and the taste, as we see, o
Page 1125 and 1126:
230. Mr. J. Jenner Weir has lately
Page 1127 and 1128:
dishments, for they nested and prod
Page 1129 and 1130:
United States and observing the bir
Page 1131 and 1132:
Turning now to domesticated and con
Page 1133 and 1134:
There is reason to believe that pig
Page 1135 and 1136:
(23. Boitard and Corbie, 'Les Pigeo
Page 1137 and 1138:
y Mr. Sclater as a distinct species
Page 1139 and 1140:
Female birds not only exert a choic
Page 1141 and 1142:
prefers the younger to the older he
Page 1143 and 1144:
ornaments the males with rare excep
Page 1145 and 1146:
differ fundamentally from ours. Fro
Page 1147 and 1148:
can forms which until lately were n
Page 1149 and 1150:
sexes, or a greater effect on one s
Page 1151 and 1152:
does not appear that intermediate g
Page 1153 and 1154:
Macgillivray, 'History of British B
Page 1155 and 1156:
to vary, both with animals in a sta
Page 1157 and 1158:
eyes have been preserved or augment
Page 1159 and 1160:
eeds of the fowl. In all the breeds
Page 1161 and 1162:
cases we have feathers symmetricall
Page 1163 and 1164:
eptiles and fishes, on the skin of
Page 1165 and 1166:
engraved from a beautiful drawing,
Page 1167 and 1168:
differently coloured fowls, and the
Page 1169 and 1170:
lue, indented centre, surrounded by
Page 1171 and 1172:
y Mr. Bartlett that they resemble t
Page 1173 and 1174:
fered fundamentally from those of t
Page 1175 and 1176:
satisfaction, this appeared to be s
Page 1177 and 1178:
cock. Many female progenitors of th
Page 1179 and 1180:
Fig.59. Portion of one of the secon
Page 1181 and 1182:
shades off downwards into a pale-le
Page 1183 and 1184:
and might easily be overlooked. The
Page 1185 and 1186:
the lower side with a fulvous tint.
Page 1187 and 1188:
tic ornament (b, Fig. 59). The uppe
Page 1189 and 1190:
parts of the ball can be perceived,
Page 1191 and 1192:
to account for the imperfect condit
Page 1193 and 1194:
dified through natural selection fo
Page 1195 and 1196:
ied on until the wonderful ball-and
Page 1197 and 1198:
feathers of humming-birds differ, w
Page 1199 and 1200:
space than all the rest of the bird
Page 1201 and 1202:
the conspicuous black colour of the
Page 1203 and 1204:
do would be to persevere in selecti
Page 1205 and 1206:
sed with the male common pheasant,
Page 1207 and 1208:
striae. So again Mr. Tegetmeier has
Page 1209 and 1210:
The same process would have to be f
Page 1211 and 1212:
form of transmission into the other
Page 1213 and 1214:
mind the reader that if sexually-li
Page 1215 and 1216:
courtship, the males alone would ac
Page 1217 and 1218:
gued that the females had aborigina
Page 1219 and 1220:
from inheriting them, on account of
Page 1221 and 1222:
tails of considerable length. The f
Page 1223 and 1224:
having been from the first transmit
Page 1225 and 1226:
contrast of colour between the sexe
Page 1227 and 1228:
themselves; that those of the secon
Page 1229 and 1230:
injured, than during cool, cloudy,
Page 1231 and 1232:
of the above forty birds the female
Page 1233 and 1234:
in equally well-concealed spots, bu
Page 1235 and 1236:
the natural selection of so-called
Page 1237 and 1238:
occasionally in a considerable degr
Page 1239 and 1240:
that here the tail differs in colou
Page 1241 and 1242:
strengthened by what Malherbe state
Page 1243 and 1244:
etween the sexes of the same specie
Page 1245 and 1246:
group will not surprise any one who
Page 1247 and 1248:
cases. In like manner the same form
Page 1249 and 1250:
feathers; and does not completely a
Page 1251 and 1252:
given by Mr. R. Sharpe, 'Proceeding
Page 1253 and 1254:
or the males alone, become extremel
Page 1255 and 1256:
of the equal transmission of charac
Page 1257 and 1258:
can be maintained with confidence,
Page 1259 and 1260:
(Emberiza) resemble one another, an
Page 1261 and 1262:
immature plumage approximately shew
Page 1263 and 1264:
I. When the adult male is more beau
Page 1265 and 1266:
it will suffice to call to mind the
Page 1267 and 1268:
ear-tufts, but the female is remark
Page 1269 and 1270:
y some remarkable facts recorded by
Page 1271 and 1272:
winter and summer, nearly the same
Page 1273 and 1274:
tions to which they will almost alw
Page 1275 and 1276:
white, whilst the female P. apoda i
Page 1277 and 1278:
with the peacock, pheasant, and fow
Page 1279 and 1280:
former period. But since this perio
Page 1281 and 1282:
tion, before given, is possible; na
Page 1283 and 1284:
protection; and it is possible that
Page 1285 and 1286:
ptarmigans, whilst changing from th
Page 1287 and 1288:
discover, of species with the femal
Page 1289 and 1290:
WHEN THE ADULT FEMALE IS MORE CONSP
Page 1291 and 1292:
the males are often kept by the nat
Page 1293 and 1294:
widely different in closely-allied
Page 1295 and 1296:
comparison is somewhat difficult to
Page 1297 and 1298:
llent account of the habits of this
Page 1299 and 1300:
teris and night-jar (Eurostopodus),
Page 1301 and 1302:
exposed to danger whilst sitting on
Page 1303 and 1304:
WHEN THE ADULT MALE RESEMBLES THE A
Page 1305 and 1306:
WHEN THE ADULT MALE RESEMBLES THE A
Page 1307 and 1308:
tection; so that they have probably
Page 1309 and 1310:
and all ages. But these characters
Page 1311 and 1312:
and the young of the sparrow of Pal
Page 1313 and 1314:
e distinguished from the female whe
Page 1315 and 1316:
placed by coloured ones. (36. Mr. B
Page 1317 and 1318:
les. But we have no reason to suppo
Page 1319 and 1320:
chives Neerlandaises,' tom. vi. 187
Page 1321 and 1322:
of N. America, Fringilla tristis, L
Page 1323 and 1324:
egard to the young differing greatl
Page 1325 and 1326:
CLASS VI. THE YOUNG IN THEIR FIRST
Page 1327 and 1328:
in the present class is that the ma
Page 1329 and 1330:
In considering this last case, if a
Page 1331 and 1332:
have occurred at different periods
Page 1333 and 1334:
confined to the females, have been
Page 1335 and 1336:
statement is too strong.) Calling t
Page 1337 and 1338:
they would have been much more cons
Page 1339 and 1340:
cailzie, blackcock, black scoter-du
Page 1341 and 1342:
Belt believes ('The Naturalist in N
Page 1343 and 1344:
tial character, for it is developed
Page 1345 and 1346:
when it discovers and flies down to
Page 1347 and 1348:
ted tail-feathers or elongated cres
Page 1349 and 1350:
progenitors of these two species, w
Page 1351 and 1352:
not endure for ever, as we may infe
Page 1353 and 1354:
fied means for producing various so
Page 1355 and 1356:
differ, the males differ much more
Page 1357 and 1358:
ornament, for producing various sou
Page 1359 and 1360:
loured, by the selection of success
Page 1361 and 1362:
iod of reproduction arrives; conseq
Page 1363 and 1364:
for having looked over these four c
Page 1365 and 1366:
ved the same fact with the hides of
Page 1367 and 1368:
dants, degenerated in size but not
Page 1369 and 1370:
jaw, or in both, are much larger in
Page 1371 and 1372:
secretion from the gland is not poi
Page 1373 and 1374:
thinner, and less branched than in
Page 1375 and 1376:
log. Generale,' 1841, p. 513. Other
Page 1377 and 1378:
the horns of the female Ant. euchor
Page 1379 and 1380:
fixed character in the females than
Page 1381 and 1382:
well as the possession of horns by
Page 1383 and 1384:
peculiar shape as in the female, bu
Page 1385 and 1386:
ce of tusks in some female walruses
Page 1387 and 1388:
"make a cut down and a jerk up, wit
Page 1389 and 1390:
[Fig. 63. Oryx leucoryx, male (from
Page 1391 and 1392:
a little backwards, he would be com
Page 1393 and 1394:
comes to the conclusion that their
Page 1395 and 1396:
[Fig. 64. Strepsiceros Kudu (from S
Page 1397 and 1398:
forward from the brow, and terminat
Page 1399 and 1400:
Male quadrupeds, which are furnishe
Page 1401 and 1402:
horns, and they rarely have canine
Page 1403 and 1404:
Tennent, 'Ceylon,' 1859, vol. ii. p
Page 1405 and 1406:
With mammals, when, as is often the
Page 1407 and 1408:
variations in strength, size, and c
Page 1409 and 1410:
half ounces, and at the age of six
Page 1411 and 1412:
Scotch deer-hound may probably be a
Page 1413 and 1414:
pons, like those of the European bo
Page 1415 and 1416:
certain extent used for this purpos
Page 1417 and 1418:
to which he is liable; for the male
Page 1419 and 1420:
Judge Caton that he had never obser
Page 1421 and 1422:
generally, or at least often, exert
Page 1423 and 1424:
of harems is nearly full. Then the
Page 1425 and 1426:
ted by Alex. Walker, 'On Intermarri
Page 1427 and 1428:
preference for each other. Finally,
Page 1429 and 1430:
strong individual antipathies and p
Page 1431 and 1432:
dies (2. Ibid. p. 595.)) of stags p
Page 1433 and 1434:
habits of other male quadrupeds. Th
Page 1435 and 1436:
History of Mamm. Animals,' 1841, p.
Page 1437 and 1438:
females. In another allied kind of
Page 1439 and 1440:
the castoreum of the beaver, see Mr
Page 1441 and 1442:
These glands secrete a semi-fluid f
Page 1443 and 1444:
sing the voice of the stag, we may
Page 1445 and 1446:
of the above-named antelopes, the P
Page 1447 and 1448:
neck, which is much less developed
Page 1449 and 1450:
that any character, even with anima
Page 1451 and 1452:
I will first give briefly all the c
Page 1453 and 1454:
Gray states, "that the males are or
Page 1455 and 1456:
With Ruminants sexual differences o
Page 1457 and 1458:
In the Indian black-buck (A. bezoar
Page 1459 and 1460:
ker than in the female; but during
Page 1461 and 1462:
nus; the young of the former, and I
Page 1463 and 1464:
male not only by his immense mane,
Page 1465 and 1466:
and differs from the young of both
Page 1467 and 1468:
ted in their transmission. Neverthe
Page 1469 and 1470:
In an earlier chapter we have seen
Page 1471 and 1472:
emasculated at an early period, los
Page 1473 and 1474:
esemblance. The hare on her form is
Page 1475 and 1476:
ed cats in his house. Such cats, as
Page 1477 and 1478:
curiously-ornamented Tragelaphus sc
Page 1479 and 1480:
guished by colour, even by the deal
Page 1481 and 1482:
the female), to species in which ne
Page 1483 and 1484:
form linking together two groups, f
Page 1485 and 1486:
stripes on the shoulders and even o
Page 1487 and 1488:
Before we conclude, it will be well
Page 1489 and 1490:
cheek. The appearance of Cercocebus
Page 1491 and 1492:
pearance and bright colours. The Se
Page 1493 and 1494:
y of Mammalia,' 1841, p. 460; see a
Page 1495 and 1496:
cases to be merely ornamental, thou
Page 1497 and 1498:
The law of the equal transmission o
Page 1499 and 1500:
mana, but not so great as in some,
Page 1501 and 1502:
character. She comes to maturity at
Page 1503 and 1504:
tions have been made in other parts
Page 1505 and 1506:
etween man and the Quadrumana, for
Page 1507 and 1508:
instance with the Aymaras and Quich
Page 1509 and 1510:
arms, in various races; and nearly
Page 1511 and 1512:
lay Arch.' vol. ii. 1869, p. 178.)
Page 1513 and 1514:
hair, and in the general shape of t
Page 1515 and 1516:
to whom they are attached; and, of
Page 1517 and 1518:
ultimately have been obliterated. T
Page 1519 and 1520:
With respect to differences of this
Page 1521 and 1522:
names under each subject, the two l
Page 1523 and 1524:
jection of Women,' 1869, p. 122), "
Page 1525 and 1526:
early education of boys and girls w
Page 1527 and 1528:
herited this difference from his ea
Page 1529 and 1530:
The sounds produced by fishes are s
Page 1531 and 1532:
aya perhaps forms an exception, as
Page 1533 and 1534:
quently be prolonged to two or thre
Page 1535 and 1536:
that an old dog of his howls when B
Page 1537 and 1538:
produced during the season of court
Page 1539 and 1540:
arts of singing and of dancing are
Page 1541 and 1542:
tion of one useful power will bring
Page 1543 and 1544:
cadences and rhythm are instinctive
Page 1545 and 1546:
clusion to that at which I have arr
Page 1547 and 1548:
love, that we have no means of judg
Page 1549 and 1550:
525-545; all the following statemen
Page 1551 and 1552:
various tints. In different countri
Page 1553 and 1554:
and Africa even the eyebrows and ey
Page 1555 and 1556:
her four front teeth from the lower
Page 1557 and 1558:
persuade some girls to give up the
Page 1559 and 1560:
maintained that savages are quite i
Page 1561 and 1562:
enormous ears"(57. Quoted by Pricha
Page 1563 and 1564:
illustrissimus viator dixit mihi pr
Page 1565 and 1566:
The Banyai of the more southern par
Page 1567 and 1568:
(62. For the Javans and Cochin-Chin
Page 1569 and 1570:
It is remarkable that throughout th
Page 1571 and 1572:
in regard to the Polynesians, in 'A
Page 1573 and 1574:
lue eyes with aversion, and they th
Page 1575 and 1576:
increasing the apparent elevation o
Page 1577 and 1578:
her barbarous nations have generall
Page 1579 and 1580:
with respect to the human body. It
Page 1581 and 1582:
CHAPTER XX. SECONDARY SEXUAL CHARAC
Page 1583 and 1584:
marry into a much lower rank. The m
Page 1585 and 1586:
observable in the erees or nobles i
Page 1587 and 1588:
hardly be added that with all savag
Page 1589 and 1590:
valuable work on 'Primitive Marriag
Page 1591 and 1592:
The indirect evidence in favour of
Page 1593 and 1594:
into strict though temporary unions
Page 1595 and 1596:
Although the manner of development
Page 1597 and 1598:
cies of baboons. (9. Brehm ('Thierl
Page 1599 and 1600:
more lax nature than that of civili
Page 1601 and 1602:
etc., tom. ii. pp. 94, 116) enters
Page 1603 and 1604:
and would not have selected the mor
Page 1605 and 1606:
offspring to inherit their beauty t
Page 1607 and 1608:
At a very early period, before man
Page 1609 and 1610:
that was possible, would be reared;
Page 1611 and 1612:
or confirming their privilege." We
Page 1613 and 1614:
any wish or expectation on the part
Page 1615 and 1616:
having been the selected. We recogn
Page 1617 and 1618:
and perhaps some other characters,
Page 1619 and 1620:
assured that no instance occurs of
Page 1621 and 1622:
The Rev. J. Shooter, 'On the Kafirs
Page 1623 and 1624:
lower animals, namely, the greater
Page 1625 and 1626:
have seen the possibility of its se
Page 1627 and 1628: has not been limited either by sex
Page 1629 and 1630: influential in leading to this kind
Page 1631 and 1632: eards equally developed in both sex
Page 1633 and 1634: y hair from their faces as somethin
Page 1635 and 1636: and with the American natives it no
Page 1637 and 1638: The colour of the face differs much
Page 1639 and 1640: that the females were modified in o
Page 1641 and 1642: nose, the colour of the skin, the l
Page 1643 and 1644: done, one path towards error is clo
Page 1645 and 1646: descendant with other mammals of a
Page 1647 and 1648: accounted for by any form of select
Page 1649 and 1650: stage in the process of modificatio
Page 1651 and 1652: the same individual, and with the m
Page 1653 and 1654: has well remarked, the largeness of
Page 1655 and 1656: and disapprobation of his fellows;
Page 1657 and 1658: happiness, praise or blame is besto
Page 1659 and 1660: not improbable that after long prac
Page 1661 and 1662: man, as Sir J. Lubbock has shewn, p
Page 1663 and 1664: what is still more important, their
Page 1665 and 1666: throughout life. In almost every gr
Page 1667 and 1668: through sexual selection in relatio
Page 1669 and 1670: individuals of one sex were during
Page 1671 and 1672: ding of the ball-and-socket ornamen
Page 1673 and 1674: clusions at which I have arrived ar
Page 1675 and 1676: in body or mind; but such hopes are
Page 1677: uted the social instincts, which af
Page 1681 and 1682: tions, which is well worthy of stud
Page 1683 and 1684: Macacus rhesus and nemestrinus) act
Page 1685 and 1686: days, "umdrehte und dem Mannchen mi
Page 1687 and 1688: the habit of congregating has been
Page 1689 and 1690: offspring of the former, although v
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The Descent of Man

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