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Science of Water : Concepts and Applications

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<strong>Water</strong> Ecology 197<br />

FIGURE 6.32 Damselfl y.<br />

leaving them to hatch or by inserting eggs singly into a slit in the stem <strong>of</strong> a submerged plant. The<br />

incomplete metamorphosis (egg, nymph, mature nymph, <strong>and</strong> adult) can take 2–3 years. Nymphs are<br />

<strong>of</strong>ten covered by algal growth.<br />

√ Note: Adult dragonfl ies are sometimes called “mosquito hawks” because they eat such a large<br />

number <strong>of</strong> mosquitoes that they catch while they are fl ying.<br />

Damselfl ies are smaller <strong>and</strong> more slender than dragonfl ies. They have three long, oar-shaped<br />

feathery tails, which are actually gills, <strong>and</strong> long slender legs (see Figure 6.32). They are gray, greenish,<br />

or brown to black in color. Their habits are similar to those <strong>of</strong> dragonfl y nymphs, <strong>and</strong> they emerge<br />

from the water as adults in the same manner. The adult damselfl ies are slow <strong>and</strong> seem uncertain<br />

in fl ight. Wings are commonly black or clear, <strong>and</strong> their bodies are <strong>of</strong>ten brilliantly colored. When<br />

at rest, they perch on vegetation with their wings closed upright. Damselfl ies mature in 1–4 years.<br />

Adults live for a few weeks or months. Unlike the dragonfl ies, adult damselfl ies rest with their wings<br />

held vertically over their backs. They are predaceous <strong>and</strong> mostly feed on live insect larvae.<br />

√ Note: Relatives <strong>of</strong> the dragonfl ies <strong>and</strong> damselfl ies are some <strong>of</strong> the most ancient <strong>of</strong> the fl ying<br />

insects. Fossils have been found <strong>of</strong> giant dragonfl ies with wingspans up to 720 mm that lived<br />

long before the dinosaurs!<br />

NONINSECT MACROINVERTEBRATES<br />

√ Note: Noninsect macroinvertebrates are important to any discussion <strong>of</strong> stream <strong>and</strong> freshwater<br />

ecology because many <strong>of</strong> them are used as bioindicators <strong>of</strong> stream quality.<br />

Three frequently encountered groups in running water systems are Oligochaeta (worms),<br />

Hirudinea (leeches), <strong>and</strong> Gastropoda (lung-breathing snails). They are by no means restricted to<br />

running water conditions <strong>and</strong> the great majority <strong>of</strong> them occupy slow-fl owing marginal habitats<br />

where the sedimentation <strong>of</strong> fi ne organic materials takes place.<br />

(1) OLIGOCHAETA (FAMILY TUIFICIDAE, GENUS Tubifex)<br />

Tubifex worms (commonly known as sludge worms) are unique because <strong>of</strong> the fact that they build<br />

tubes. Sometimes there are as many as 8000 individuals/m 2 . They attach themselves within the tube<br />

<strong>and</strong> wave their posterior end in the water to circulate the water <strong>and</strong> make more oxygen available

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