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Annona Species Monograph.pdf - Crops for the Future

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Chapter 9. Genetic Improvement<br />

crosses, were planted. Since attractiveness is a key factor in selling fruits, <strong>the</strong><br />

exciting possibility exists to develop new cultivars with external and internal<br />

pink-red colours. Crossing of newly introduced red and pink-skinned<br />

atemoya x cherimoya selections to red-skinned sugar apple types selected in<br />

Queensland, Australia, is currently in progress (George et al., 1999).<br />

Spain is more focused on germplasm collection and ex situ evaluation of<br />

numerous accessions (Farré Massip and Hermoso González, 1987, cited by<br />

Scheldeman, 2002). Pérez de Oteyza and Farré (1999) reported that <strong>the</strong><br />

selection of a superior cultivar of cherimoya at <strong>the</strong> Experimental Station of<br />

La Mayora, Spain, is based on <strong>the</strong> following characters: a) regarding<br />

agronomic and commercial parameters - size, <strong>for</strong>mat and pilosity on leaves,<br />

length and colour of flowers, floral density (number of flowers per one metre<br />

of mature stem), susceptibility to fruit fly, season of maturation and harvest,<br />

defects on <strong>the</strong> skin and in <strong>the</strong> pulp, resistance to fungal attack and seed<br />

colour; and b) regarding fruit transport and consumption - type of skin, pulp<br />

firmness, taste (sugar content and acidity) and seed index (number of seeds<br />

per 100 g of fruit).<br />

In Chile, introduction and selection of cherimoya showed that <strong>the</strong> new<br />

cultivars from Spain were superior, with longer harvest periods and better<br />

fruit quality than Chilean cultivars (Gardiazabal and Cano, 1999). Ten<br />

cultivars from Spain were evaluated <strong>for</strong> fruit weight, shape and colour, skin<br />

type, number of days post-harvest to reach appropriate ripeness <strong>for</strong> eating,<br />

resistance of pulp to pressure, percentage by weight of fruit components,<br />

seed type, number of seeds per 100 g of pulp, sugar and acidity, and taste.<br />

The Spanish cultivar ‘Cholan’ showed <strong>the</strong> highest general rating.<br />

In Mexico, genetic improvement of cherimoya started in 1991, with<br />

evaluation of seedlings of local cultivars and evaluation of introduced<br />

cultivars from Spain, Chile and New Zealand (Román and Damián, 1999).<br />

Characterisation and selection of cherimoya fruits from trees collected in<br />

three regions of Michoacán State, Mexico, was done by Agustín (1999).<br />

While studying such characteristics as fruit weight, percentage of pulp and<br />

seed, fruit soundness, type of skin and earliness, he found great genetic<br />

variability among native fruits and proposed this as <strong>the</strong> basis <strong>for</strong> developing<br />

germplasm <strong>for</strong> commercial plantations. A very similar strategy has been used<br />

in Italy and Portugal (Madeira), with selection and evaluation of promising<br />

local types and introduced cultivars (Monastra, 1997; Nunes, 1997).<br />

The methods and strategies used in Madeira to develop superior cultivars of<br />

cherimoya were somewhat different. First, <strong>the</strong> agricultural service surveyed<br />

63

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