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Annona Species Monograph.pdf - Crops for the Future

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Chapter 9. Genetic Improvement<br />

and mass selected local types. Plant behaviour was observed with respect to<br />

age, origin, farming practices, pest and disease resistance, organoleptic<br />

parameters and so on. At <strong>the</strong> same time, genetic material was introduced<br />

from o<strong>the</strong>r countries, especially <strong>the</strong> USA and Spain, increasing variability<br />

and allowing comparison of adaptability, productivity and quality with <strong>the</strong><br />

local materials. Finally, clonal selection produced four improved cultivars:<br />

Madeira, Mateus I, Perry Vidal, and Funchal (Nunes, 1997).<br />

Soursop is still largely propagated by seed, and <strong>the</strong> progenies can be selected<br />

and separated into groups based on acidity and sweetness. Morada, an<br />

ecotype introduced into Brazil from Colombia, produces large fruits with an<br />

average weight of 3.8 kg, but its fruits have very acid pulp, which is a<br />

constraint <strong>for</strong> <strong>the</strong> fresh fruit market. A local soursop selection from Bahia<br />

State named "ecotype A" produces very small fruits (< 1.8 kg), but has sweet<br />

pulp, appropriate <strong>for</strong> <strong>the</strong> fresh market. Crossing between <strong>the</strong>se two might<br />

produce a medium weight fruit with a commercially acceptable taste. Several<br />

genotypes and some related species are used in <strong>the</strong> soursop breeding work of<br />

Embrapa Cerrados, Brazil, from which a clone based on Morada has been<br />

released as a reliable option <strong>for</strong> tolerance to trunk borer (Pinto and Ramos,<br />

1997). At <strong>the</strong> same time, A. glabra, which has dwarf characteristics and good<br />

adaptation to damp areas, is being used in a breeding programme <strong>for</strong><br />

rootstock improvement, since <strong>the</strong>re is acceptable rootstock/scion<br />

compatibility between <strong>the</strong>se two species.<br />

Wild soursop has been used <strong>for</strong> insect control in Africa, since it has<br />

significant amounts of secondary metabolites with insecticidal activity (see<br />

Chapter 7). However, <strong>the</strong>re is no specific breeding programme to improve<br />

this characteristic, although Abubakar and Abdurahman (1998) mentioned a<br />

project to prospect, collect and chemically identify wild soursop variation<br />

with insect growth-regulating activity in Kaduma State, Nigeria (work at <strong>the</strong><br />

Department of Biological Sciences, Ahmadu Bello University, Zaria).<br />

9.4.4 Selection and cultivar development<br />

Like any o<strong>the</strong>r fruit tree, <strong>the</strong> type of selection or cultivar development of<br />

annonas is strongly influenced by commercial use. Also, <strong>the</strong> feasibility of<br />

using a particular type of selection or cultivar may depend on <strong>the</strong> nature of<br />

<strong>the</strong> organisations that produce and distribute grafted plants <strong>for</strong> commercial<br />

use, <strong>the</strong>se being much better established in more developed countries. This<br />

explains why annona cultivar development concentrates on cherimoya and<br />

soursop (and atemoya), and has been carried out only by public institutions.<br />

64

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