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Physiology and Molecular Biology of Stress ... - KHAM PHA MOI

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188<br />

K. Janardhan Reddy<br />

constitution <strong>of</strong> the plant, chemical constituents <strong>of</strong> the soil, climatic conditions <strong>and</strong> age<br />

<strong>of</strong> the plant. Despite wide variation in the mineral composition <strong>of</strong> different plants, a<br />

certain critical level <strong>of</strong> nutrients is necessary for healthy growth <strong>of</strong> plants. An idea <strong>of</strong><br />

elemental composition with relative levels <strong>of</strong> each <strong>of</strong> the nutrients for higher plants is<br />

indicated in Table 1.<br />

Table 1. Concentration <strong>of</strong> mineral elements in soil, available form <strong>and</strong> their<br />

content in plants*<br />

Element Available form Content in soil Content in dry<br />

matter (µmol/g)<br />

Molybdenum<br />

2-<br />

MoO 4<br />

0.2 – 2.0 ppm 0.001<br />

Nickel Ni 2+ < 100 ppm 0.001<br />

Copper Cu + & Cu 2+ 5 – 50 ppm 0.10<br />

Zinc Zn 2+ 10 – 30 ppm 0.30<br />

Manganese Mn 2+ 200 – 300 ppm 1.00<br />

Iron Fe 2+ & Fe 3+ 2,50,000 ppm 2.00<br />

Boron BO 3<br />

20 – 200 ppm 2.00<br />

Chlorine Cl - 3.00<br />

Sulphur<br />

2-<br />

SO 4<br />

0.04% 30<br />

Phosphorus H 2<br />

PO 4- 0.1% 60<br />

Magnesium Mg 2+ 0.05% 80<br />

Calcium Ca 2+ 0.5% 125<br />

Potassium K + 2.0% 250<br />

Nitrogen<br />

+<br />

NO 3-<br />

& NH 4<br />

1000<br />

*Adapted from Epstein (1965), Brown et al. (1987), Hopkins <strong>and</strong> Hüner (2004).<br />

Nutrient stress is a complex phenomenon, underst<strong>and</strong>ing <strong>of</strong> which requires<br />

the co-ordinated efforts <strong>of</strong> soil scientists, ecologists, physiologists, biochemists,<br />

agronomists <strong>and</strong> molecular biologists. Nutrient stress may result from either by low<br />

levels <strong>of</strong> availability <strong>of</strong> the element or by the presence <strong>of</strong> excess concentrations. In<br />

some cases the presence <strong>of</strong> one element in excess concentrations may induce the<br />

deficiency <strong>of</strong> another element. In this context, attempts were made to show the availability,<br />

functional aspects, <strong>and</strong> deficiency <strong>and</strong> toxicity symptoms <strong>of</strong> 14 elements essential for<br />

the survival <strong>of</strong> plants <strong>and</strong> other elements, which produce phytotoxicity. Visual deficiency<br />

symptoms provide a valuable basis for assessing the nutritional status <strong>of</strong> the plant.<br />

Deficiency symptoms are the consequence <strong>of</strong> metabolic disturbances at various stages

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