ILLUSTRATED FLORA OF EAST TEXAS - Brit - Botanical Research ...
ILLUSTRATED FLORA OF EAST TEXAS - Brit - Botanical Research ...
ILLUSTRATED FLORA OF EAST TEXAS - Brit - Botanical Research ...
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CURRENT VEGETATION <strong>OF</strong> PINEYWOODS/INTRODUCTION 95<br />
Pityopsis graminifolia (narrow-leaf silk-grass), Solidago odora (anise-scented goldenrod),<br />
Tephrosia virginica (goat’s-rue), and various species of Panicum (panic grasses). Also present are<br />
indicators of dry sandy sites, including Pteridium aquilinum (bracken fern), Tragia urticifolia<br />
(nose-burn), Cnidoscolus texanus (Texas bull-nettle), Stylisma pickeringii (Pickering’s dawnflower),<br />
and Berlandiera pumila (soft greeneyes). Recurring fires keep shrubs sparse and<br />
maintain the grassy ground layer. With the exception of longleaf pine, whose seedlings are<br />
adapted to surviving fire, trees have difficulty becoming established in regularly burned<br />
stands. Rare on modern landscapes, these communities persist mainly on public lands in<br />
areas with prescribed burning programs. Good examples are found in the southern Angelina<br />
National Forest.<br />
LOAMY DRY-MESIC UPLANDS—This ecological type has sandy loam or loam surface soils with<br />
loam or clay loam more than 30 cm below the surface. Consequently, these soils hold more<br />
moisture than Arenic Dry Uplands. They represent the least-droughty examples of the “dry<br />
uplands” of Nixon (2000), and the “dry upland forests and savannas” of Harcombe et al.<br />
(1993). On most Pineywoods landscapes they are the most widespread upland ecological<br />
type. As with Arenic Dry Uplands, we recognize two natural plant communities, mixed<br />
pine-hardwood forests and longleaf pine woodlands, depending on whether stands have<br />
had a history of regular fire and are within the range of longleaf pine.<br />
DRY-MESIC MIXED PINE-HARDWOOD UPLANDS—On these uplands (Fig. 54), Pinus taeda (loblolly<br />
pine) is abundant, even dominant, but usually occurs with a mixture of broad-leaved<br />
deciduous trees, including Liquidambar styraciflua (sweetgum), Quercus falcata (southern<br />
red oak), Q. stellata (post oak), Ulmus alata (winged elm), and Carya spp. (hickory). Shortleaf<br />
pine (Pinus echinata) may also be present, especially on sites of more natural quality, although<br />
human activities and lack of fire have favored loblolly pine and have reduced the occurrence<br />
of shortleaf pine even in the northern and western Pineywoods where it was always an<br />
important component of Dry-Mesic Uplands. The shrub-layer may be dense and includes<br />
Callicarpa americana (American beauty-berry), Ilex vomitoria (yaupon), and Cornus florida<br />
(flowering dogwood). Woody vines, especially Smilax spp. (greenbrier) and Vitis aestivalis<br />
(summer grape) are common. A variety of species with a wide ecological range, including<br />
among others, Chasmanthium sessiliflorum (narrow-leaved wood-oats), Toxicodendron radicans<br />
(poison ivy), and Parthenocissus quinquefolia (Virginia creeper) dominate the ground layer.<br />
These mixed forest communities have increased markedly on the post-settlement landscape<br />
at the expense of longleaf pine woodlands.<br />
DRY-MESIC LONGLEAF PINE UPLANDS—These stands, which correspond in part with the “longleaf<br />
bluestem uplands” of Ajilvsgi (1979), are typically open-canopied Pinus palustris (longleaf<br />
pine) woodlands, often with scattered individuals of Quercus marilandica (blackjack oak).<br />
The sparse canopy allows enough light to reach the ground to support a dense, prairie-like<br />
ground layer of grasses, composites, and other sun-loving species. Schizachyrium scoparium<br />
(little bluestem) usually dominates the ground layer. Other important species include<br />
Pityopsis graminifolia (narrow-leaf silk-grass), Solidago odora (anise-scented goldenrod),<br />
Tephrosia virginica (goat’s-rue), Panicum virgatum (switch grass), and Dichanthelium (rosette<br />
grass) species. Frequent fire controls shrubs such as Rhus copallina (flame-leaf sumac), Ilex<br />
vomitoria (yaupon), and Liquidambar styraciflua (sweetgum) saplings, which rapidly invade<br />
unburned sites. While longleaf pine seedlings are capable of surviving fire, regular fire prevents<br />
seedlings of most other species from becoming established. In the absence of regular<br />
fire, sites rapidly succeed to a mixed closed-canopy forest. More favorable soil moisture and<br />
nutrient conditions allow fire-free stands to convert to mixed forests more quickly than<br />
would Arenic Longleaf Pine Uplands. Historically, these Dry-Mesic Longleaf Pine Uplands<br />
dominated uplands in much of the southern and eastern Pineywoods. They are now rare,