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ILLUSTRATED FLORA OF EAST TEXAS - Brit - Botanical Research ...

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118 INTRODUCTION/VEGETATION <strong>OF</strong> POST OAK SAVANNAH<br />

Americans were present and possibly had significant impacts on the vegetation through such<br />

mechanisms as increased fire frequency. Holloway et al. (1987), based on pollen analysis of<br />

Weakly Bog in Leon County, suggested that the Post Oak Savannah became established only<br />

between 1,500 and 2,000 years ago as increasing aridity resulted in a vegetational shift from<br />

oak woodland to savannah-like plant communities. While the pollen data may represent a<br />

significant one-time shift in the vegetation, there have been numerous changes in vegetation<br />

since the end of the last glaciation (Bousman 1998), and the continuing “state of flux”<br />

(Smeins 1984) mentioned above should not be forgotten. Because of all these complexities,<br />

the exact presettlement vegetation pattern is particularly difficult to estimate.<br />

Animal life on the Post Oak Savannah was in general a mixture of that found on the<br />

adjacent Blackland Prairies and Pineywoods. Since detailed discussions are given for the<br />

Blackland Prairie and Big Thicket portion of the Pineywoods, no further discussion of presettlement<br />

animal life is given here.<br />

As settlers moved into the area, much of the Post Oak Savannah was rapidly converted<br />

to cropland or pasture. In the words of David Diamond of the Texas Natural Heritage<br />

Program (in Bartlett 1995):<br />

The prairies are more fertile, yet initially were harder to plow because they were clay soils, and so<br />

they were not plowed as quickly as the Post Oak Savanna. A lot of the savannas were plowed. They<br />

grew cotton or other crops, and the soil wore out quickly because it wasn’t as fertile as the Blackland<br />

Prairies. Later on, when the prairies were plowed, the Post Oak Savanna went back into rangeland<br />

of some sort and today is no longer plowed.<br />

Today, much has been cleared and large areas are occupied by economically useful exotics.<br />

As pointed out by Simpson (1988), it is “used largely for ‘improved’ pasture, with vast<br />

acreages sodded to Bermuda Grass and seeded to Bahia Grass.” On the other hand, large<br />

expanses of once-cleared land are now in second, third, or fourth growth woody vegetation.<br />

Thus, as a result of human impacts, little of the Post Oak Savannah is as it was during<br />

presettlement times.<br />

VEGETATION <strong>OF</strong> THE POST OAK SAVANNAH<br />

As mentioned above, while probably dominated originally to a significant extent by savannahs/open<br />

woodlands/forests of post oak and blackjack oak with a tall grass understory, the<br />

Post Oak Savannah was a complex mosaic of plant communities. The communities listed<br />

below, modified in part from Wilson (1989), Bezanson (2000), and MacRoberts et al.<br />

(2002b), represent the primary plant communities and give some indication of the complexity<br />

of the vegetation of the Post Oak Savannah. It should be noted, however, that there were and<br />

still are small areas of different, and in some cases, quite distinctive communities. An excellent<br />

example of this can be seen at the Gus Engeling Wildlife Management Area in Anderson<br />

Co., where Singhurst et al. (2003b) recently documented a number of interesting “vegetation<br />

alliances,” some of which have similarities to communities of the Pineywoods. These include<br />

the Sand Post Oak-Bluejack Oak Alliance (Quercus margaretta-Q. incana), the White Oak-<br />

Southern Red Oak-Post Oak Alliance (Quercus alba-Q. falcata-Q. stellata), the Loblolly Pine-<br />

Southern Red Oak Alliance (Pinus taeda-Quercus falcata), the Overcup Oak Seasonally<br />

Flooded Forest Alliance (Q. lyrata), and the Planer Tree Seasonally Flooded Forest Alliance<br />

(Planera aquatica).<br />

POST OAK-BLACKJACK OAK UPLAND SAVANNAHS, WOODLANDS, AND FORESTS—The dominant natural<br />

vegetation for much of the upland Post Oak Savannah was an open deciduous<br />

savannah/woodland/forest composed of Quercus stellata (post oak), Q. marilandica (blackjack<br />

oak), and other drought-tolerant species (e.g., Carya texana, black hickory) (Bezanson 2000)<br />

and an understory of tall grasses such as Schizachyrium scoparium (little bluestem),

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