ILLUSTRATED FLORA OF EAST TEXAS - Brit - Botanical Research ...
ILLUSTRATED FLORA OF EAST TEXAS - Brit - Botanical Research ...
ILLUSTRATED FLORA OF EAST TEXAS - Brit - Botanical Research ...
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218 INTRODUCTION/ORIGIN AND DIVERSITY <strong>OF</strong> <strong>EAST</strong> <strong>TEXAS</strong> <strong>FLORA</strong><br />
FIG. 129/MAP SHOWING SURFACE EXTENT <strong>OF</strong> THE CARRIZO SANDS.ONLY 19 KM (12 MILES) WIDE, THE CARRIZO FORMATION TRENDS SOUTH-<br />
WEST TO NORTH<strong>EAST</strong> FOR 720 KM (450 MILES) (FROM SORRIE & WEAKLEY 2001, WITH PERMISSION <strong>OF</strong> THE SOUTHERN APPALACHIAN<br />
BOTANICAL SOCIETY).<br />
Thelesperma flavodiscum (East Texas greenthread) (Sorrie & Weakley 2001).<br />
A different pattern of West Gulf Coastal Plain endemism (and one with fewer examples)<br />
is shown by those species that occur only on the Weches Formation (thin rocky soils with<br />
high pH and glauconite). Two such East Texas endemics are Texas golden glade cress<br />
(Leavenworthia aurea var. texana) (Mahler 1987; Poole et al. 2002) and the federally endangered<br />
white bladderpod (Lesquerella pallida) (George & Nixon 1990).<br />
EDWARDS PLATEAU ENDEMICS—Many endemics that were once thought to be restricted to the<br />
Edwards Plateau are now known to reach the westernmost edge of East Texas. Additionally,<br />
many of the Edwards Plateau endemics also occur in the Lampasas Cut Plain in the southern<br />
part of the Cross Timbers and Prairies vegetational area (Amos & Gehlbach 1988; Diggs et<br />
al. 1999). The explanation for the endemism seen in the Edwards Plateau and adjacent areas,<br />
while not completely clear, may be the result of the climatic history of the last 1.8 million<br />
years. During the Quaternary Period (beginning about 1.8 mya), there were significant climatic<br />
variability and at least 20 glacial-interglacial cycles. Widespread changes in vegetation<br />
were associated with these climatic fluctuations (Delcourt & Delcourt 1993). For example,<br />
during the last full-glacial interval (100,000 until approximately 18,000–15,000 years ago),<br />
there was a cool, moist “pluvial” climate across the unglaciated parts of southwestern North<br />
America (Delcourt & Delcourt 1993) with forest species presumably expanding their ranges.<br />
Bryant’s data (1977) indicated an open deciduous forest in central Texas during the last fullglacial<br />
interval. The climate moderated from 15,000–10,000 years ago, with interglacial conditions<br />
(i.e., warmer and drier) for the last 10,000 years (Delcourt & Delcourt 1993), (e.g.,<br />
Nolina lindheimeriana (devil’s-shoestring), Tinantia anomala (false dayflower, Tradescantia<br />
edwardsiana (plateau spidewort), and Yucca rupicola (twist-leaf yucca)). The Edwards Plateau<br />
endemics are typically found in moist areas such as canyons along wooded streams and have<br />
presumably survived in the favorable microclimate pockets as the overall climate of the area<br />
has warmed and/or dried. Many of these species have affinities with eastern taxa and may be<br />
relicts of a more widespread flora that became restricted as the result of climatic or geologic<br />
changes (Palmer 1920; Amos & Gehlbach 1988).