ILLUSTRATED FLORA OF EAST TEXAS - Brit - Botanical Research ...
ILLUSTRATED FLORA OF EAST TEXAS - Brit - Botanical Research ...
ILLUSTRATED FLORA OF EAST TEXAS - Brit - Botanical Research ...
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136 INTRODUCTION/PRESETTLEMENT AND EARLY SETTLEMENT CONDITIONS ON BLACKLAND PRAIRIE<br />
Grayson County] its name, the trees being a major landmark in a featureless terrain.”<br />
(McLeRoy 1993). The riverine forests along Big Mineral Creek (Grayson County) were<br />
described by Parker (1856) as “a rich bottom, thickly grown up with large cotton wood,<br />
honey locust, overcup [actually bur oak], and other heavy timber, besides plenty of the bois<br />
d’arc.” Maclura pomifera (bois d’arc) was apparently endemic to a small area in northeast TX<br />
(12 counties, mostly in the Blackland Prairie) and adjacent OK and AR (Little 1971; Weniger<br />
1996). Based on early explorer accounts (e.g., Marryat 1843), this species sometimes formed<br />
impenetrable thickets along creeks in the northern Blackland Prairie (see detailed discussion<br />
in Weniger 1996). Roemer (1849) described a trading post he visited in Falls County as “on<br />
a hill covered with oak trees, two miles distant from the Brazos, above the broad forested bottom<br />
of Tohawacony Creek.” He further described the wooded bottomland as having “high, dense<br />
trees.” The presence of such wooded bottomlands was probably quite striking in the midst of<br />
vast stretches of prairie.<br />
In summary, the original vegetation of the Blackland Prairie seems to have been predominantly<br />
tall grass prairie with trees along watercourses, sometimes scattered on the<br />
prairie, or concentrated in certain areas (e.g., Pilot Grove), possibly as the result of locally<br />
favorable soil conditions or topography.<br />
THE ROLE <strong>OF</strong> FIRE ON THE PRESETTLEMENT BLACKLAND PRAIRIE—Fire was probably an important (if<br />
not the most important) factor in the maintenance of the presettlement Blackland Prairie vegetation,<br />
and it undoubtedly had a major impact on the structure of plant communities<br />
(Anderson 1990; Collins & Gibson 1990; Strickland & Fox 1993; Jones & Cushman 2004).<br />
Axelrod (1985) argued that fire was a primary factor in the rise of the extensive North<br />
American grasslands after the end of the last glacial maximum. Unlike woody plants, which<br />
are usually killed or severely damaged by fire, grasses are highly fire-adapted, with a number<br />
of specific adaptations allowing them not only to survive but prosper under conditions of<br />
recurrent fire (and also grazing and trampling). These adaptations include the presence of<br />
intercalary meristems (located in the culms just above the nodes and in the leaves near the<br />
ligules, thus allowing growth from the base even if terminal parts are damaged), the large<br />
amount of below ground biomass, and the tendency to branch (“tiller”) or produce stolons<br />
or rhizomes near or below ground level.<br />
During early settlement times, huge grassland fires were well known in central North<br />
America—for example, a massive 1885 fire described by Haley (1929) began in western<br />
Kansas and burned across northern Texas, a total distance of 282 km (175 miles). Haley<br />
(1929) also gave many other examples of large-scale grass fires. He quoted Hank Smith, an<br />
early settler in Crosby County in the Texas plains country, about a fire in 1879:<br />
The fire swept thousands of square miles of country to the south and southwest, north and northeast<br />
of Mount Blanco. All through the country at that time, especially along the streams were hundreds of<br />
magnificent groves of fine timber, particularly cottonwood and hackberry.… That fire killed the<br />
timber and in effect literally wiped it out.<br />
Another example is a 1905 fire that started at Hyannis, Nebraska, presumably ignited by<br />
lightning. “It traveled over 165 miles and finally burned out within the watersheds of the<br />
Middle Loup and Dismal Rivers, apparently stopped by both rivers” (Komarek 1966).<br />
The Blackland Prairie, located at the extreme southern end of the True Prairie grassland<br />
association (Gould & Shaw 1983), would appear to have been especially susceptible<br />
to prairie fires. The high summer temperatures, extremely irregular summer rainfall,<br />
periodic droughts, strong winds, frequent summer lightning storms, and resulting recurrent<br />
fires during hot dry periods would have been potent forces in shaping the vegetation