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564 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIX oblongo-lanceolata, integerrima, penninervia; pedunculi tenues, bini, profunde bifidi, ramo altero florifero, altero cirrhoso, pedunculo altero simplici, ecirrhoso; bracteae 3, liberae, foliaceae, anguste ovatae; calycis tubus patelliformis; sepala elliptico-ovata, aristata; petala sepalis paullo breviora; coronae filamenta pauca, linearia, 1-seriata; staminum filamenta antheris breviora; ovarium anguste ellipsoideum. Herbaceous vine, glabrous throughout; stem slender, terete, striate; tendrils slender, of 2 forms, one axillary on the main stem, the other on one of each pair of peduncles; stipules setaceous, 6 to 8 mm. long, subpersistent; petioles 8 to 10 mm. long, slender, canaliculate on the upper side, biglandular just below the apex, the glands sessile, 1 to 1.5 mm. in diameter; leaves oblong or oblong-lanceolate, 4 to 11 cm. long, 1.5 to 5 cm. wide, cuspidate-acuminate, subrotund at base, entire, obscurely glandular at margin, penninerved (secondary nerves about 9 to a side), conspicuously reticulate, coriaceous, lustrous on both surfaces, drying light green; peduncles slender, 3.5 to 7 cm. long, 1-flowered, in pairs, one simple, ecirrhose, the other bifid, with 1 branch floriferous and the other a tendril; bracts 3, free to the base, narrowly ovate, 2 to 3 cm. long, 0.5 to 1 cm. wide, obtuse at the apex, acute and sessile at the base, conspicuously reticulate, sparingly and obtusely glandular at the margin, chartaceous, sublustrous; calyx tube very short, barely 1 mm. long, about 3 mm. wide; sepals elliptic-ovate, about 1.5 cm. long, 3 to 6 mm. wide, obtuse, 3-nerved (lateral nerves obscure), the midnerve terminating dorsally in a filiform awn 2.5 to 3 mm. long; petals slightly shorter and narrower than the sepals, obtuse, very thin; corona consisting of a few linear filaments 2 to 3 mm. long, apparently in a single series; filaments of stamens 3 mm. long, the anthers 6 to 7 mm. long; gynophore about 6 mm. long; ovary narrowly ellipsoidal. Type in the Herbarium of Field Museum of Natural History, No. 622,608, collected at Tarapoto, Department of Loreto, Peru, altitude 750 meters, December, 1929, by Llewelyn Williams (No. 5902). Duplicate in the United States National Herbarium (No. 1,470,181). Peru. DISTRIBUTION: Known only from the type locality, in northern Quite possibly this curious plant belongs to an undescribed genus of Passifloraceae, but the flowers are young and most of them are in such poor condition that it is impossible to determine the nature or position of the genital organs, and I am therefore reluctant to base a
AMERICAN PASSIFLOBACEAE 565 new genus upon it. There appear to be five stamens, with thick filaments, borne directly upon the calyx, and three styles. One flower, with a definite gynophore, shows no vestiges of stamens; another, with five protuberances on the floor of the calyx, which may be the remnants of stamens, lacks a gynophore. It is impossible to say whether this is due to some of the parts having become detached or whether the flowers are unisexual, like those of many species of the Old World genus Adenia. From all members of Passiflora this is at once differentiated by the tendrils and peduncles. The large bracts are like those of certain granadillas, notably P. nitida and P. riparia, though of a rather different texture. It is to be hoped that, as a result of the intensive botanical exploration of northern Peru, plants with more fully developed flowers will be collected. DOUBTFUL SPECIES Passiflora maculata Scan, in Colla, Hort. Ripul. 101. 1824. This was described from a cultivated plant of unknown origin. De Candolle gave a slightly amplified description in the Prodromus (p. 324), adding "In Curacao. Pluk. t. 210, f. 3. An P. minima Lin.(?) v. s." The Plukenet figure, accompanied by the description "Flos <strong>Passion</strong>is Curassavicus . . . ," almost certainly represents P. suberosa, though no petiolar glands are shown. This inclusion of the Plukenet reference is apparently the only reason for ascribing P. maculata to Curasao The specimen labeled P. maculata in the . De Candolle Herbarium, probably type material, is well preserved, and clearly does not belong to any of the species treated in the present monograph. Its vegetative parts are exceedingly well matched, strange to say, by a specimen, without flowers, in the United States National Herbarium, bearing the data "Sea beach, Brunswick Co., N. C., Aug. 5, 1885, G. McCarthy." From the De Candolle specimen I have drawn up the following description: Herbaceous vine; stem angulate, sparingly pilose or glabrescent; stipules setaceous, about 5 mm. long; petioles up to 1.5 cm. long, glandless; leaves 3-lobed about four-fifths their length (upper ones less deeply lobed ; lobes oblong-lanceolate, 5 to 6 cm. long, 1 to 2 cm. wide, the lateral often divergent at nearly a right angle from the middle lobe), subcoriaceous, bright green and glabrous above, paler and pilosulous on the nerves beneath, white-maculate; peduncles subequal to the petioles; bracts setaceous; flowers about 1.5 cm. wide;
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THE UNIVERSITY OF ILLINOIS
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THE AMERICAN SPECIES OF PASSIFLORAC
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PRINTED IN THE UNITED STATES OF AME
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UNIVERSITY OF ILLINOIS-URBANA
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