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386 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIX (BM). Cuyaba, Lindman A3483 (S); Malme 1220B (S); Pilger 184 (B), 218 (B); Moore 917 (B, BM), 961 (B, BM, Y). Minas Geraes: Claussen 382 (BM, Gen, P, S). Rio de Janeiro: Carandahy, Glaziou 12740 (B, Cop, Gen, P). Sao Paulo: St. Hilaire 753 (P). Santa Catharina: Ule 1669 (P). PARAGUAY: Itagua, Hassler 1332 (Bo, P, Y). Rio Apa, Hassler 8130 (A, B, BM, Bo, type of P. cincinnata var. imbricata, Gen, P, V); Fiebrig 4804 (B, BM, Gen, K). Cordillera de Altos, Hassler 3339 (B, Bo, Gen, P, V). Bahia, Anisits 2271 (S). Asuncion, cultivated, Lindman A1755 (S). La Trinidad, Balansa 2204, in part (Bo). ARGENTINA: Salta: Tartagal, Venturi 7629 (N). Formosa: Jorgensen 2617 (G, Mo, N); Meyer 945 (N). These specimens show much variation in the shape of the leaves, and it now seems impossible to maintain as a separate species the Venezuelan plant described as P. perlobata. This handsome vine we found well established along the trail between Chinacota and La Esmeralda, in eastern Colombia. LOCAL NAMES: "Pachis" (Bolivia); "maracuja," "tubarao" (Brazil). 229. Passiflora filamentosa Cav. Diss. 10: 461. pi. 294. 1790. (t)Passiflora cuneifolia Cav. Diss. 10: 460. pi. 292. 1790. Plant glabrous or minutely villosulous; stem terete; stipules dolabriform, 3 to 4 mm. long, 1 to 1.5 mm. wide, aristate, serrulate on one side, very soon deciduous; petioles up to 4 cm. long, biglandular above middle, the glands sessile or subsessile; leaves 4 to 8 cm. long, 6 to 12 cm. wide at greatest width, normally palmately 5-lobed about four-fifths their length (occasionally one or both of the lowest lobes merged into the adjacent lobe so that the leaf is 4-3-lobed, the lobes oblong-lanceolate, 1 to 3 cm. wide, acuminate, serrulate, usually minutely glandular in the sinuses), cordulate at base, membranous, sublustrous above, glabrous or minutely villosulous; peduncles up to 5 cm. long; bracts ovate or ovate-spatulate, 1 to 2 cm. long, 8 to 12 mm. wide, rounded or acutish at apex, narrowed at base, serrulate except often in lower half; flowers up to 8 cm. wide; calyx tube campanulate; sepals narrowly oblong, 8 to 10 mm. wide, obtuse, white, green-keeled without, the keel terminating in a subulate awn about 3 mm. long; petals subequal to or slightly longer than the sepals, white; corona filaments filiform, in several series, those of the 2 outer series radiate, longer than to half as long as the sepals,
AMERICAN PASSIFLORACEAE 387 banded with blue and white, the filaments of the inner series densely massed, erect, 2 to 5 mm. long, blue; operculum membranous in lower half, filamentose in upper half; nectar ring annular; limen cupuliform; ovary ovoid, tomentulous; fruit globose, about 4 cm. in diameter, very acid, the pericarp brittle; seeds obcordate, about 10 mm. long, 7 mm. wide, strongly flattened, broadly wing-margined, punctate at center. TYPE LOCALITY: "America," the type seen at Madrid. ILLUSTRATIONS: Cav. Diss. 10: pi. 294; Bot. Mag. 46: pi. 2023; Bot. Reg. 7: pi. 584- DISTRIBUTION: Probably central Brazil. BRAZIL: Minas Geraes: Serra da Gramma, 1,000 meters, Chase 9548 (N). The taxonomic position of this species is doubtful. The broadly winged seeds at once suggest a close relationship with P. quadrangularis, but in no other detail are the two species alike. The shape and serrulation of the leaves are strongly suggestive of P. serratodigitata, but the bracts, more like those of P. edulis, and the coronal arrangement are quite different. Masters placed the species near P. violacea and P. amethystine,, in the group with large, foliaceous, subreniform, persistent stipules, but the stipules are very small and early deciduous. In the present treatment P. filamentosa is $aced near P. cincinnata, though it differs in several important details. Perhaps it is a hybrid between P. caerulea and P. serrato-digitata. Passiflora filamentosa has been frequently cultivated in Europe, but, as Masters observed, specimens of positively native origin are very rarely found in herbaria. The type, in the Cavanilles Herbarium, is in a fine state of preservation, agreeing perfectly with Mrs. Chase's specimen from Minas Geraes, which was collected in a very dense jungle, with every indication that the plant was native. Regarding the origin of the type specimen Cavanilles says, "In America. Colui in horto Issy. Exc. Ducis ab Infantado." Passiflora cuneifolia, considered by Masters as possibly identical with P. edulis, I take to be a form of P. filamentosa with 3-lobed leaves, though perhaps it is a hybrid between P. filamentosa and P. caerulea. 230. Passiflora recurva Mast, in Mart. Fl. Bras. 13, pt. 1: 608. 1872. Plant glabrous throughout; stem subangular; stipules linearsubulate, about 3 mm. long, soon deciduous; petioles 1 to 2.5 cm.
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THE UNIVERSITY OF ILLINOIS
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THE AMERICAN SPECIES OF PASSIFLORAC
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PRINTED IN THE UNITED STATES OF AME
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UNIVERSITY OF ILLINOIS-URBANA
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