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350 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIX L. Williams 4980 (N); Poeppig 2171 (V, type of P. colubrina}. Rio Ucayali, Tessmann 3155 (B). Junin: San Ramon, cultivated, Kittip & Smith 24092 (F, N, Y). BOLIVIA: Beni: Junction of Beni and Madre de Dios rivers, Rusby 490 (B, BM, Bo, G, K, N, Ph, Y). Bopi River, Mulford Biol Expl. 739 (Y). La Paz: Mapiri, Buchtien 1676 (N). Coroico, Buchtien 3851 (N). Santa Cruz: Santa Cruz, Herzog 1507 (B, S). Rio Surutu, Steinbach 7220 (BM, G, Gen, Ph, S, Ut). The leaves of P. triloba are usually 3-lobed, minutely serrulatedenticulate, and generally larger than in its near relatives. The species is sometimes confused with P. ligularis and P. tiliaefolia. discussion of the differences between these three species found under P. tiliaefolia. 202. Passiflora tiliaefolia L. Sp. PI. 956. 1753. A will be Plant glabrous throughout; stem terete; stipules ovate-lanceolate to oblong-lanceolate, 1 to 2 cm. long, 0.5 to 1.5 cm. wide, or the upper linear-lanceolate and barely 2 mm. wide, acute or acuminate, oblique at base, entire or serrulate, green, often turning reddish in drying; petioles 2.5 to 7 cm. long, 2^i-glandular, the glands saucershaped, all borne at apex or 1 pair near middle, subsessile or shortstipitate, 1 to 2 mm. wide; leaves cordate-ovate, 10 to 25 cm. long, 8 to 18 cm. wide, abruptly acuminate, shallowly or deeply cordate at base (basal lobes not overlapping), entire, membranous, often lustrous on both surfaces; peduncles 2 to 3 cm. long; bracts ovate, about 2 cm. long, 1.5 cm. wide (perhaps larger), obtuse or acute at apex, united one-fifth to one-third their length, reddish when dry, glabrous except for a narrow band of tomentum along the margin within; flowers about 8 cm. wide; sepals oblong, 5 to 7 mm. wide, obtuse, concave, slightly carinate on outside, the keel terminating in a slender awn; petals oblong, subequal to sepals; corona about 5-ranked, the two outer rows of filaments terete, about half as long as petals, the three inner rows barely 3 mm. long; operculum membranous, incurved, entire at margin; limen adnate to gynophore; ovary ovoid; fruit ovoid, about 6 cm. in diameter, "deep violet to blackish proximally, greenish distally." TYPE LOCALITY: Lima, Peru. ILLUSTRATIONS: Journ. Obs. Phys. Math. Bot. Peruv. 2: pi. 121. 1714; Amoen. Acad. 1: pi. 10, /. 4; Cav. Diss. 10: pi. 285; G. Don, Hist. Dichl. PI. 3: 51. /. 4; Mutis, Icon. PI. Ined. 26: pi. 6.
AMERICAN PASSIFLORACEAE 351 DISTRIBUTION: Mountains of Colombia and Peru, 1,500 to 2,500 meters altitude. COLOMBIA: Mutis 794 (Ma, N), 2219 (Ma, N), 2275 (Ma, N), 2890 (Ma, N). Tolima: Above Ibague", Killip & Hazen 9625 (G, Ma, Ph, Y). Antioquia: Medellin, Toro 214 (Y). Caldas: Salento, Killip & Hazen 9027 (B, G, N, Ph, Y), 9078 (N). Pereira, Killip & Hazen 11006 (N). El Valle: La Cumbre, KiUip 5697 (N). El Cauca: Cuatro Esquinas, Pennell & Killip 6350 (N). PERU: Pav6n (B, BM, Gen, Ma). The exact identity of P. tiliaefolia is by no means clear. Apparently both Linnaeus and Cavanilles knew the plant only from the description and figure given by Feuille"e. This description in general would apply almost equally well to either P. ligularis, entire-leaved forms of P. triloba, or the specimens cited above. thus: The differences between these three species may be summarized P. ligularis Stipules ovate-lanceolate or oblong-lanceolate. Petiolar glands filiform or narrowly liguliform, 3 mm. long or more. Leaves usually not lobed, entire at margin. Basal lobes not overlapping. Bracts less than 4 cm. long, united one-fifth to one-third. Corona 5-ranked, the 2 outer ranks subequal to petals, the 3 inner ranks much shorter. Operculum denticulate. P. triloba Stipules ovate-lanceolate. Glands saucer-shaped, stipitate, less than 3 mm. long. Leaves usually 3-lobed, minutely denticulate. Basal lobes overlapping. Bracts more than 4 cm. long, free or united only at very base. Corona 3-ranked, the outermost rank half as long as petals, the second rank minutely denticulate. Operculum crenulate. P. tiliaefolia Stipules ovate-lanceolate or oblong-lanceolate, or the upper narrowly linear-lanceolate. l Glands saucer-shaped, subsessile or short-stipitate, less than 3 mm. long. Leaves never (or rarely?) lobed, entire at margin. Basal lobes overlapping. Bracts less than 4 cm. long, united one-fifth to one-half. Corona 5-ranked, the 2 outer ranks about half as long as petals, the 3 inner ranks much shorter. Operculum entire. Plate 12 is lacking from the copy of FeuilleVs work which I consulted ; I assume that Cavanilles' illustration arid the figure of a leaf in the Amoenitates are substantially accurate reproductions. In both of 1 In the Pav6n plant of P. tiliaefolia in the British Museum the lower stipules are ovate-lanceolate, about 3 cm. long, 1.5 cm. wide, the upper linear-lanceolate, 1 cm. long, 0.4 cm. wide. The specimen of this same collection at Berlin has linearlanceolate stipules, 2 cm. long, 0.2 cm. wide. In the Colombian material the stipules are uniformly ovate-lanceolate.
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Synonyms in italics. INDEX Page num
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UNIVERSITY OF ILLINOIS-URBANA
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