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570 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIX near the base of the ovary, free, or united close to the base; anthers linear-oblong, 2-celled, versatile; gynophore erect; ovary 1-celled, with 4 parietal placentae; styles 4, distinct to the base; stigmas reniform-capitate; fruit ovoid. TYPE SPECIES: Mitostemma Glaziovii Mast. REFERENCE: Gontscharow, Bull. Jard. Princ. U. S. S. R. 26: 556-558. 1927. Styles much longer than the ovary (British Guiana). Styles shorter than the ovary (Brazil). Ovary glabrous; leaves sharply acuminate at apex. 1. M. Jenmanii. 2. M. Glaziovii. Ovary white- villous; leaves obtuse or obtusely acuminate at apex. 3. M. brevifilis. Our knowledge of this genus is still altogether too imperfect, especially in regard to the shape of the corona filaments. The only adequate specimen in the United States National Herbarium is one of Glaziovii, and from that the description of the corona given above M . has been derived. The descriptions of the corona presented by Masters, Harms, and Gontscharow differ from each other in certain details. Masters amplified his Latin diagnosis of the corona thus: "The corona springs from the mouth of the very short flower-tube, and consists of a large number of separate, threadlike, thick processes arranged in a triple series; the outermost are terete, acute, fleshy, reddish orange, somewhat shorter than the petals; next to these is a series of lobes like those just described, but each has a membranous, lacerate wing on either side, so that the thick, fleshy thread is, as it were, a midrib between the two membranous wings; the third and innermost series consists of a number of oblong processes, wholly membranous, crisped, and lacerate at the edges." I believe the coronal structure is similar in all three species. Mitostemma is even more like Dilkea than has been supposed; the presence of a well defined calyx tube to distinguish Dilkea cannot be relied upon, inasmuch as the sepals at length separate, leaving a very small calyx, which, as in Mitostemma, is not much more than the enlargement of the top of the peduncle. 1. Mitostemma Jenmanii Mast. Journ. Bot. Brit. & For. 21: 34. 1883. Scandent shrub(?), glabrous throughout; tendrils few, stout; branches terete or subangular; petioles about 1 cm. long, suberose,
AMERICAN PASSIFLORACEAE 571 glandless; leaves oblong or oblong-obovate, 6 to 9 cm. long, 3 to 4.5 cm. wide, abruptly narrowed to an obtuse apex, rounded or acutish at base, entire, penninerved (principal lateral nerves 8 to 10 to a side), reticulate-veined with scarcely elevated veins, thick-coriaceous, sublustrous above, dull beneath; flowers about 4 cm. wide, in short, axillary racemes, the rachis scarcely 5 mm. long, the pedicels 5 to 8 mm. long, articulate near base; bracts setaceous, 1 to 2 mm. long, 2 borne at point of articulation, the third near base of peduncle; calyx tube short-campanulate; sepals oblong, about 2 cm. long, 0.6 cm. wide, obtuse, reddish orange; petals similar to and slightly shorter than the sepals; outermost filaments of the corona 8 to 10 mm. long, reddish orange, the inner ones slightly shorter, white; stamens 1 to 1.3 cm. long; ovary fusiform; styles longer than the ovary. TYPE LOCALITY: Mazaruni River, British Guiana. DISTRIBUTION: Known only from the type locality. BRITISH GUIANA: Mazaruni River, Jenman 622 (EG, K, type). 2. Mitostemma Glaziovii Mast. Journ. Bot. Brit. & For. 21^34. 1883. Dilkea Glaziovii Mast, ex Glaziou, Bull. Soc. Bot. France 56, Me"m. 3:313. 1909. Scandent shrub, glabrous throughout; tendrils few, stout; stem terete, purplish; stipules filiform, about 4 mm. long; petioles 1 to 1.5 cm. long, stout, glandless; leaves oblong, 6 to 18 cm. long, 3.5 to 7 cm. wide, acute or abruptly acuminate, rounded at base, entire, penninerved (principal lateral nerves 9 to 11 to a side), conspicuously reticulate-veined, coriaceous, lustrous, without ocellae or occasionally with 2 glands at base; peduncles solitary, 2.5 to 4 cm. long, articulate below middle, slightly thickened toward apex; bracts setaceous, one about 3 mm. long, borne near base of peduncle, 2 half as long, at point of articulation; calyx tube short-campanulate, 8 to 10 mm. wide, introrse, green; sepals and petals similar and subequal, oblong, about 1.5 cm. long, 5 to 6 mm. wide, obtuse, white; outermost corona filaments reddish orange, about 7 mm. long, the others about 5 mm. long, white; stamens about 1 cm. long; gynophore 5 to 7 mm. long; ovary ellipsoidal, acute; fruit broadly ovoid, 2 cm. long or more, acutish. TYPE LOCALITY: Near Rio de Janeiro, Brazil. ILLUSTRATIONS: Engl. & Prantl, Pflanzenfam. 3, 6a: 72. /. 25; ed.2,21:477./.2&T.
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THE UNIVERSITY OF ILLINOIS
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THE AMERICAN SPECIES OF PASSIFLORAC
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PRINTED IN THE UNITED STATES OF AME
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