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572 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIX DISTRIBUTION: Vicinity of Rio de Janeiro, Brazil. BRAZIL: Rio de Janeiro: Lagoa dos Peixes, near Cabo Frio, Glaziou 12741 (Gen, K, type, P); Widgren 1165 (Cop, S). Corcovado, Glaziou 6089 (Cop, P, S); Nadeaud in 1862 (P). Mundo Novo, Kuhlmann 15328 (Bo, K, N, S, Ut). 3. Mitostemma brevifilis Gontsch. Bull. Jard. Bot. Princ. U. S. S. R. 26: 557. 1927. Shrub 90 to 120 cm. high, the branchlets terete, glabrous; tendrils present; petioles 3 to 4.5 mm. long, glandless; leaves oblong-lanceolate, lanceolate, or oblanceolate, 5 to 14 cm. long, 3.5 to 5.5 cm. wide, obtuse or obtusely acuminate, rarely emarginate, narrowed at base, subrevolute, penninerved, coriaceous, glabrous; flowers solitary (or in racemes?); bracts linear-setiform, 2 to 3.5 mm. long; calyx tube broadly campanulate, very short; sepals 4, oblong, 1.5 to 1.7 cm. long, 6 mm. wide, obtuse, sericeo-puberulent, white; petals 4, similar to and slightly narrower than the sepals, white; corona "blue," the 2 outer series 8.5 to 9 mm. long, the innermost about 5 mm. long; stamens 8, the filaments about 8 mm. long, blue; ovary ellipticoblong, white- villous; styles shorter than the ovary; fruit subglobose, 5 to 5.5 cm. long, about 3 cm. in diameter, densely velutinous. TYPE LOCALITY: Rio Pardo, southern Brazil. DISTRIBUTION: Known only from the type locality. BRAZIL: Rio Grande do Sul: Rio Pardo, Riedel 535 (K, type collection). I am not certain that this inflorescence should be described as racemose. In the only specimen which I have seen the lower flowers are solitary in the axils of the leaves. Perhaps these racemes are floriferous branches bearing much reduced, caducous leaves. 4. DILKEA Mast. Trans. Linn. Soc. 27: 627. 1871. Woody vines, subscandent shrubs, or small trees, without tendrils or rarely with a few poorly developed ones; leaves alternate, petiolate, simple, penninerved; bracts minute, subulate, borne near base of pedicels; flowers in axillary or terminal glomerules or short-spicate, rarely solitary, hermaphrodite, red or white; sepals 4 (or 5?), oblong, fleshy, united below middle when young to form a cylindric or funnelshaped tube, at length separating to the base and readily deciduous, leaving a very small receptacle; petals 4 or 5, slender, free to the base; corona 2-ranked, the outer rank consisting of slender, free or nearly
AMERICAN PASSIFLORACEAE 573 free filaments, the inner tubular in the lower part, cleft above into floccose, crispate threads or into segments margined with such threads; operculum none; stamens 8, equal, hypogynous, free except at the very base; anthers linear or linear-oblong, versatile; ovary subsessile or short-stipitate, with 4 parietal placentae; styles 4, united below the middle, exserted; stigmas reniform-capitate; fruit globose or ovoid, the pericarp coriaceous; seeds 8 to 10, large, with a parchment-like covering. TYPE SPECIES: Dilkea retusa Mast. This genus apparently is confined to the little explored recesses of the Amazon basin, and our knowledge of it is still too imperfect to permit of more than a tentative treatment of the species. Although the genus itself is a well marked one, readily recognized among Passifloraceae by the conspicuous fringe of pale, crispate hairs on the second series of corona filaments, the species are for the most part poorly defined. Dr. Ducke, who after many years of travel in the upper Amazon region is more familiar with the living plants than anyone else, has recently made the interesting suggestion to me 7 that Dilkea is a monotypic genus. I am inclined to think that further study and collecting will show that the four last species of the present treatment should be merged in one; the key to these, it will be observed, is a weak one. Dilkea parviflora, however, seems clearly distinct. In describing Dilkea Masters treated two species, D. retusa and D. acuminata, and in the Flora Brasiliensis he added a third one, D. Wallisii. A fourth species was described by Barbosa Rodriguez in 1891, and a fifth one is proposed in the present paper. According to Masters' illustration in the Flora Brasiliensis (pi. 106), P. retusa, P. acuminata, and P. Wallisii appear to have distinctive leaf outlines, but Ducke has observed (Archiv. Jard. Bot. Rio de Janeiro 3: 222. 1922; 5: 174. 1930) that D. Wallisii has heteromorphic foliage, and his No. 21307 shows this. Masters' illustrations show rather prominent glands near the middle of the petioles, but in neither his generic nor his specific descriptions does he mention the presence of petiolar glands. In the material I have examined, including the types of two of Masters' species, glands normally are not present, though occasionally one or more of the petioles may bear a pair of scarlike glands. Masters' description of the flower tube as cylindric, with the petals borne at its throat and the corona near its middle, is not substantiated by the specimens I have at hand. At first the sepals are
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THE UNIVERSITY OF ILLINOIS
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THE AMERICAN SPECIES OF PASSIFLORAC
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PRINTED IN THE UNITED STATES OF AME
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