downloads/Killip 2.pdf - Passion Flowers

566 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIX
petals none; corona filaments apparently in a single series; operculum
membranous, plicate(?).
2. TETRASTYLIS Barb. Rodr. Rev. Engenharia 4: 260. 1882.
Passiflora Sect. Tetrastylis Harms, in Engl. & Prantl, Pflanzenfam.
1. Aufl. 1. Nachtr. 256. 1897.
Woody or herbaceous vines, bearing simple, axillary tendrils;
stipules present; leaves alternate, petiolate; flowers in axillary racemes,
or solitary or in pairs in the axils of the leaves; calyx tube
short, patelliform; sepals 5; petals 5, alternate with the sepals,
inserted at margin of tube; corona filamentose; operculum membranous;
gynophore elongate, curved; stamens 5, the filaments
monadelphous, united beyond the gynophore into a broad membrane,
united at
only the tips free; anthers oblong, bifid at base; styles 4,
very base; ovary oblong, stipitate, obtusely 4-angled; ovules on 4
parietal placentae.
TYPE SPECIES: Tetrastylis montana Barb. Rodr.
The genus Tetrastylis was established by Barbosa Rodriguez in
1882, and to it was assigned a single Brazilian species, Tetrastylis
montana Barb. Rodr. The description of the plant was very complete
and was accompanied by an excellent illustration. The principal
points of difference between this genus and Passiflora, as noted
by Barbosa, were: Tetrastylis: four styles; gynophore curved; stamen
filaments united beyond gynophore, only the extremities free; four
placentae; Passiflora: three styles; gynophore straight; stamen
filaments free from gynophore to extremities; three placentae.
In the first edition of the Naturlichen Pflanzenfamilien Harms
recognized Tetrastylis as a valid genus, placing it immediately before
Passiflora. In the supplement to this work he created the section
Tetrastylis, in Passiflora, for this species, a course followed in the
second edition.
REFERENCE: Killip, Tetrastylis, a genus of Passifloraceae, Journ.
Wash. Acad. Sci. 16: 365-369. 1926.
Flowers in axillary racemes; leaves entire, coriaceous; stipules
filiform,
soon deciduous; petioles glandular at base; woody vine
(Brazil) 1. T. ovalis.
Flowers solitary or in pairs in the axils of the leaves; leaves 3-lobed,
membranous; stipules semi-ovate, persistent; petioles glandular
at middle; herbaceous vine (Costa Rica) 2. T. lobata.