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396 FIELD MUSEUM OF NATURAL HISTORY BOTANY, VOL. XIX bracts, several species and varieties have been described. Masters reduced all to four varieties but admitted that they freely intergraded. Among the specimens cited above two well marked forms may be observed: in one the filaments of the two outer ranks of the corona are nearly as long as the sepals, the succeeding filaments being dentiform in two distinct rows, and the corona entire; in the other the outer two rows of filaments are less than half as long as the sepals, the succeeding filaments linear-falcate in three closely approximate series, the operculum fimbrillate at the margin. The first form is well represented by Lindman 825, the second by Chase 8250. Differences in vegetative characters, however, do not appear to be correlated with these differences in coronal structure, either in P. edulis or in P. caerulea, another commonly cultivated species exhibiting the same differences in the length of the two outer ranks of the corona. As many of the herbarium specimens doubtless represent cultivated plants or escapes, it is possible that one of the forms has resulted from the introduction of a strain of some other species. For the present I prefer not to assign formal names to these variants. Passiflora verrucifera was described as having glands on the sepals near the margin. Otherwise it appears to differ little from the form of P. edulis with proportionately longer corona filaments. No specimens that I have examined have these glands, and it is possible that the type of P. verrucifera was a hybrid between P. edulis and P. setacea. Two species described by Barbosa Rodriguez, P. picroderma and P. iodocarpa, closely resemble P. edulis in vegetative characters. The coronal structure of neither corresponds exactly to that of the two forms mentioned above. Possibly the coronal structure of this species is even more variable than here noted, or perhaps Barbosa's plants are actually distinct species. The types of these apparently are not in existence. Passiflora vernicosa Barb. Rodr. is the form of P. edulis with proportionately long outer corona filaments. Passiflora edulis is cultivated in Florida and the West Indies and in South America from Venezuela to Brazil. In many places it has escaped and become well established. In Australia it is the most commonly cultivated species of Passiflora, and its fruit is highly esteemed. It is interesting to note that the late Dr. H. K. W. Kumm, who was engaged in the cultivation of passion-fruit in southern California, developed a fourth generation plant of P. edulis with four placentae. LOCAL NAMES: "Parche" (Venezuela); "maracuja," "maracujamirim," "maracuja dedoce," "maracuja peroba" (Brazil).
AMERICAN PASSIFLORACEAE 397 Series 10. Palmatisectae 234. Passiflora palmatisecta Mast, in Mart. Fl. Bras. 13, pt. 1: 564. 1872. Stem subangular, cano-hirsutulous or the older portions glabrescent; stipules linear-subulate, 4 to 6 mm. long, deciduous; petioles up to 2 cm. long, glandless; leaves membranous, hispidulous above, cano-hirsutulous beneath, polymorphic, the lower subhastately 3-lobed to below middle, the lobes narrowly oblong, entire, dentate, or often irregularly lobulate, the middle lobe up to 5 cm. long, 1 to 2.5 cm. wide, the lateral lobes half as long; upper leaves 3-parted nearly to base, the segments 2-3-lobed nearly to their base, the lobes linear, less than 5 mm. wide, obtuse, subentire, denticulate, or lobulate-dentate; peduncles up to 2 cm. long; bracts ovate, 2 to 3 mm. long, 1 to 1.5 mm. wide, remotely ciliate-denticulate, hispidulous, borne 2 to 5 mm. below base of flower; flowers 3.5 to verticillate, 4 cm. wide; calyx tube campanulate; sepals ovate-lanceolate, about 10 mm. long, 5 mm. wide, obtuse, dorsally carinate (keel terminating in a short horn), pale green without, white within; petals linearoblong, slightly longer than the sepals, about 3 mm. wide, obtuse, white; corona in 2 ranks, the filaments narrowly liguliform, equaling the petals, pale green; operculum membranous, about 2 mm. long, horizontal, white, entire at margin; limen cupuliform, about ^ mm. high, erect, closely surrounding base of gynophore, the margin spreading; ovary ovoid, glabrous; fruit ovoid, 2.5 to 3 cm. long, about 1.5 cm. wide, 6-angled, tapering at both ends, stipitate, glabrous; seeds cuneate-oblong, about 6 mm. long, 2.5 mm. wide, subtridentate at apex, truncate at base, finely reticulate. TYPE LOCALITY: Rio Saladella de San Jago, Tucuman, Argentina. DISTRIBUTION: Southern Bolivia and northwestern Argentina, 550 to 700 meters altitude. BOLIVIA: El Chaco: Rio Pilcomayo, Fries 1568 (N, S). ARGENTINA: Salta: Cerro Negro, Venturi 10294 (N). Catamarca: San Miguel, Hieronymus & Lorentz (B). Tucuman: Burroyacu, 750 meters, Stuckert 12571 (Gen). Presito, 750 meters, Venturi 7522 (F, N), 7555 (N), 7694 (BM, G, K, N). Santiago del Estero: Cerro del Remate, Venturi 6043 (N). Cordoba: Nabor, Hieronymus & Niederlein 136 (B). La Rioja: Hieronymus & Niederlein 144 (B). The excellent material obtained by Fries and by Venturi permits a better conception of this species than heretofore has been possible.
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THE UNIVERSITY OF ILLINOIS
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THE AMERICAN SPECIES OF PASSIFLORAC
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PRINTED IN THE UNITED STATES OF AME
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UNIVERSITY OF ILLINOIS-URBANA
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