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RA 00048.pdf - OAR@ICRISAT

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to be produced and the nature of facilities that are<br />

developed. The capital investment is usually much<br />

higher in the market oriented economy because<br />

the shortage and high cost of human resources<br />

demand mechanization and automation. The finance<br />

may be received as a government grant,<br />

loan from institutions responsible for rural development<br />

and/or from commercial banks.<br />

Research Responsibility<br />

Agro-research scientists working in the universities,<br />

plant breeding stations and international<br />

research centers cooperate to maintain a continuous<br />

flow of superior hybrids and improved<br />

varieties to replace the existing ones. The superiority<br />

may be for yield, resistance to pests or<br />

diseases, ability to withstand environmental<br />

stress or any other trait of economic value. In<br />

advanced countries, many private seed enterprises<br />

have also created research facilities and<br />

thus share the responsibility to evolve better<br />

sorghum hybrids and varieties. They do not<br />

disclose their pedigree and thus enjoy a monopoly<br />

on the production and sale of their seed.<br />

In most developing countries, the responsibility<br />

to develop, test and release superior hybrids or<br />

varieties is entirely that of national research<br />

institutes. International research organizations<br />

such as ICRISAT also supply improved variety/<br />

hybrid trials and assist developing countries to<br />

plant and identify superior adaptable types. These<br />

are of open pedigree and any agency in the public<br />

or private sector can produce their seeds.<br />

Seed Classes—Their<br />

Maintenance and Increase<br />

The objective of seed production is to build a<br />

strong chain of multiplication of the various seed<br />

classes. Three stages of multiplication recognized<br />

in seed certification are:<br />

1. Breeder's seed<br />

2. Basic (foundation) seed, and<br />

3. Commercial (certified) seed.<br />

Breeder's seed is the first multiplication stage<br />

after a new superior variety or hybrid has been<br />

officially released, notified and recommended for<br />

cultivation. The responsibility for the increase of<br />

this seed class is usually with the originating<br />

breeder or his station. If the variety/hybrid has<br />

been developed through the cooperative efforts<br />

of a national or international research project, the<br />

coordinator nominates the breeder or breeders<br />

who will be responsible for breeder seed multiplication<br />

and supply within each country. In case of a<br />

hybrid, the male sterile line (A line), the maintainer<br />

line (B line) and the restorer line (R line) comprise<br />

the breeder's seed.<br />

Breeder's seed of maintainer line, restorer line<br />

and variety are multiplied in a small plot, either<br />

under complete isolation or by controlled pollination<br />

usually on a university or research farm. The<br />

male-sterile line is increased by planting in an<br />

isolated plot A and B lines in alternate sets of rows<br />

(frequently 4 : 2 or 6: 2 A and B lines respectively).<br />

Each plant is examined at the vegetative, flowering<br />

and seed ripening stages for trueness-to-type<br />

and seed-borne diseases. The breeder and his<br />

staff ensure that the seed meets the highest<br />

standards of genetic purity, as one off-type plant<br />

will give rise to hundreds of such plants in the<br />

subsequent multiplication stages. Maximum care<br />

is taken to avoid all chances of mechanical<br />

mixtures during harvesting, threshing, cleaning,<br />

grading, packing and labeling of this seed class.<br />

The general standards and specific requirements<br />

of breeder's seed are frequently not<br />

stipulated by most official seed certification agencies<br />

in the world, possibly because the authority<br />

competent to label breeder's seed is the originating/nominated<br />

breeder himself and he is beyond<br />

the purview of the certification agency. However,<br />

the breeder's seed must be field tested before it<br />

is released as a source for basic seed multiplication.<br />

The field test may be jointly conducted by the<br />

breeder and the seed certification agency.<br />

As a matter of policy and convenience, adequate<br />

quantities of breeder's seed to last for 3 - 4<br />

years should be multiplied at one time. The extra<br />

seed can be retained as a buffer stock for any<br />

sudden rise in demand or increased projected<br />

requirements. It will also avoid errors that tend to<br />

arise with frequent handling. Variations which<br />

tend to develop from delayed segregation, mutation,<br />

outcrossing, genetic drift or selection bias of<br />

the breeder will also be minimized. A suitable cold<br />

storage facility should be created at the research<br />

station to store the planned carry-over stock.<br />

Basic (foundation) seed is frequently produced<br />

from breeder's seed. It is not necessary that<br />

production of all basic seed be from breeder's<br />

seed. As long as trueness-to-type is maintained it<br />

can be produced from existing basic seed. Gener-<br />

501

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