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Yoshida - 1981 - Fundamentals of Rice Crop Science

Yoshida - 1981 - Fundamentals of Rice Crop Science

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CLIMATIC ENVIRONMENT AND ITS INFLUENCE 91<br />

Table 2.13. Heat <strong>of</strong> combustion for different parts <strong>of</strong> the rice<br />

plant . a<br />

Plant part<br />

Heat <strong>of</strong> combustion (cal/g)<br />

Leaf blade<br />

Leaf sheath<br />

Root<br />

Panicle<br />

Dead leaf<br />

Whole plant<br />

4036<br />

3569<br />

31 77<br />

3978<br />

3611<br />

3718<br />

a Mean values calculated from Murata et al (1968).<br />

go in our attempts to increase crop yields. Several scientists used different methods<br />

<strong>of</strong> estimating maximum potential rice yields using different assumptions (Evans<br />

1972, Murata and Matsushima 1975, IRRI 1977).<br />

An estimate <strong>of</strong> the maximum potential rice yield is given by the product <strong>of</strong> the<br />

maximum photosynthetic net production per day or per unit <strong>of</strong> incident solar<br />

radiation and the number <strong>of</strong> effective days for grain production.<br />

There are two basic methods for estimating maximum photosynthetic net<br />

production. Method 1 estimates the gross production by examining the efficiencies<br />

<strong>of</strong> component processes in the energy flow <strong>of</strong> canopy photosynthesis and subtracts<br />

the respiration losses to obtain the net production (Murata and Matsushima 1975).<br />

Method 2 is more empirical and much simpler. It uses the recorded maximum<br />

photosynthetic efficiency <strong>of</strong> rice crops (IRRI 1977). Both methods implicitly<br />

assume that the sink size is not limiting to grain production and that all the<br />

photosynthates are carbohydrate and contribute to grain carbohydrate.<br />

Method 1<br />

Assumptions:<br />

(a) The period for yield production is 40 days after heading.<br />

(b) The average daily solar radiation is 400 cal/cm 2 per day <strong>of</strong> which 45% is<br />

photosynthetically active.<br />

(c) Of the photosynthetically active radiation, 5.5% is lost by reflection at the<br />

canopy surface and another 10% through absorption by inactive tissues.<br />

(d) Eight photons are required to reduce 1 molecule <strong>of</strong> carbon dioxide, corresponding<br />

to an efficiency <strong>of</strong> 26% in energy conversion,<br />

(e) The loss due to light saturation in the upper leaves is 17% at 400 cal/cm 2 per<br />

day.<br />

(f) The conversion factor for dry matter is 3,900 cal/g.<br />

(g) The respiration loss is the sum <strong>of</strong> 1.5% <strong>of</strong> dry weight/day (1.5 kg/m 2 in this<br />

case) and 25% <strong>of</strong> the gross photosynthesis.<br />

(h) The ratio <strong>of</strong> husk weight to the dry grain weight is 20%.

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