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Yoshida - 1981 - Fundamentals of Rice Crop Science

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52 FUNDAMENTALS OF RICE CROP SCIENCE<br />

Table 1.13. Duration <strong>of</strong> each development stage for a panicle<br />

or a spikelet. a<br />

Developmental stages b<br />

Duration (days)<br />

1953 c 1954 d<br />

Differentiating stage <strong>of</strong> the neck node 2.0 2.0<br />

Increasing <strong>of</strong> bracts stage 1.0 1.0<br />

Duration <strong>of</strong> differentiation<br />

<strong>of</strong> bracts 3.0 3.0<br />

Early differentiating stage <strong>of</strong><br />

primary rachis-branches 1.5 1.0<br />

Middle differentiating stage <strong>of</strong><br />

primary rachis-branches 2.0 2.0<br />

Late differentiating stage <strong>of</strong><br />

primary rachis-branches 2.2 1.0<br />

Duration <strong>of</strong> differentiating stage<br />

<strong>of</strong> primary rachis-branches 5.7 4.0<br />

Early differentiating stage <strong>of</strong><br />

secondary rachis-branches 1.2 1.0<br />

Late differentiating stage <strong>of</strong><br />

secondary rachis-branches 2.0 1.5<br />

Duration <strong>of</strong> differentiating <strong>of</strong><br />

secondary rachis-branches 3.2 2.5<br />

Initial stage <strong>of</strong> spikelet<br />

differentiation 1.0 1.0<br />

Early differentiating stage<br />

<strong>of</strong> spikelets 3.0 3.0<br />

Middle differentiating stage<br />

<strong>of</strong> spikelets 1.0 1.2<br />

Late differentiating stage<br />

<strong>of</strong> spikelets 4.0 3.2<br />

Duration <strong>of</strong> differentiation<br />

<strong>of</strong> spikelets 9.0 8.4<br />

Differentiating stage <strong>of</strong> pollen<br />

mother cell 2.5 1.8<br />

Reduction division stage 2.0 1.8<br />

Early stage <strong>of</strong> extine formation 1.2 1.6<br />

Extine formation stage 3.3 2.5<br />

Early ripe stage <strong>of</strong> pollen 2.0 2.0<br />

Ripe stage <strong>of</strong> pollen 5.5 5.0<br />

Total 37.4 32.6<br />

a Matsushima (1970). b The topmost spikelets were used. c Unusual<br />

weather. d Normal weather.<br />

For example, when a 14-leaf and a 18-leaf variety have developed 6 leaves on the<br />

main culm, their leaf number indexes are 43 and 33, respectively.<br />

The leaf number indexes ate related to the developmental stages <strong>of</strong> a growing<br />

panicle (Table 1.14). Such relationship holds only for terminal spikelets and for<br />

16-leaf varieties. The neck-node differentiation occurs when the leaf number<br />

index is around 77, or when the plant has developed 12.3 leaves. The spikelet<br />

differentiation stage comes at a leaf number index <strong>of</strong> around 90, or when the plant<br />

has 14.4 leaves.

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