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Yoshida - 1981 - Fundamentals of Rice Crop Science

Yoshida - 1981 - Fundamentals of Rice Crop Science

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GROWTH AND DEVELOPMENT OF THE RICE PLANT 31<br />

1.28. Synchronous growth <strong>of</strong> a tiller, a leaf, and roots <strong>of</strong> a rice plant (adapted from Fujii 1974).<br />

After germination, one radicle emerges and elongates to a maximum length <strong>of</strong><br />

about 15 cm; it functions until the seventh-leaf stage. Mesocotyl roots emerge from<br />

the axis between the node <strong>of</strong> the coleoptile and the base <strong>of</strong> the radicle. Mesocotyl<br />

roots develop only under such conditions as deep seeding or when the seed is<br />

treated with chemicals.<br />

The rice root system is basically composed <strong>of</strong> nodal roots. Each node usually<br />

develops about 5–25 roots. When the development <strong>of</strong> root, leaf, and tiller is<br />

examined, it is more convenient to regard an internode rather than a node as a unit<br />

(Fig. 1.30). A leaf and thin roots emerge from the upper region <strong>of</strong> the internode<br />

whereas a tiller and thick roots develop from the internode’s lower region. Roots<br />

developing directly from the culm’s nodal regions are called primary roots. The<br />

diameter <strong>of</strong> the primary roots ranges from 0.5 mm to greater than 1 mm. As<br />

growth advances, the primary roots develop branched secondary roots, which in turn<br />

develop tertiary roots, and so on. Under lowland conditions, rice is known to<br />

develop sixth-order branched roots. The root diameter becomes successively

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