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Yoshida - 1981 - Fundamentals of Rice Crop Science

Yoshida - 1981 - Fundamentals of Rice Crop Science

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32 FUNDAMENTALS OF RICE CROP SCIENCE<br />

1.29. Three kinds <strong>of</strong> rice roots<br />

(Hoshikawa 1975).<br />

1.30. Structural unit <strong>of</strong> root, leaf, and<br />

tiller (Kawata et a1 1963).<br />

smaller (ranging from 1,000 to 40 µm) as the order <strong>of</strong> branching increases.<br />

The formation <strong>of</strong> root hairs is greatly affected by the root environment. Aerobic<br />

conditions in upland soils favor root hair formation (Kawata and Ishihara 1959,<br />

1961, Kawata et a1 1964); the reductive conditions <strong>of</strong> flooded soils impair it. <strong>Rice</strong><br />

root hairs are about 5–10 µm in diameter and about 50–200 µm long. Most rice<br />

varieties reach a maximum rooting depth <strong>of</strong> 1 m or deeper in s<strong>of</strong>t upland soils. In an<br />

anaerobic environment, such as flooded soils with impaired water percolation, rice<br />

roots seldom exceed a maximum depth <strong>of</strong> about 40 cm. Actual root depth found in<br />

the field is controlled not only by genetic ability but also by environmental<br />

conditions. Total root length increases as the shoot grows. At heading, it is about<br />

2–3 km/plant when plants are grown singly in a root box. In an upland field, the<br />

total root length ranges from 15 to 34 km/m 2 <strong>of</strong> soil surface. Specific root length<br />

varies from 10 to 1,000 m/g dry weight. Thick roots are shorter per unit dry weight<br />

than thin roots. The average length <strong>of</strong> rice roots is about 300 m/g dry weight, which<br />

corresponds to a root with a diameter <strong>of</strong> 0.2 mm.

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