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Yoshida - 1981 - Fundamentals of Rice Crop Science

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214 FUNDAMENTALS OF RICE CROP SCIENCE<br />

Table 6.1. Response <strong>of</strong> a traditional tall variety to graded doses<br />

<strong>of</strong> nitrogen, with and without farmyard manure (FYM). a<br />

0 33.6 67.2 100.8 134.40<br />

kg/ha kg/ha kg/ha kg/ha<br />

No manure 2.08 2.59 2.49 2.49 2.22<br />

Farmyard manure 2.44 2.72 2.47 2.17 1.92<br />

at 8.89 t/ha<br />

a Chandler (1963). b Added as ammonium sulfate. Converted from<br />

Ib/acre.<br />

Rough rice yield (t/ha) at nitrogen doses b <strong>of</strong><br />

mal water and weed control at moderately wide spacing.<br />

High yielding japonica rices have short, narrow, erect, dark-green leaves; thin,<br />

but short and sturdy culms; and short, dense panicles. They respond to increased<br />

application <strong>of</strong> nitrogen, and produce high yields (Jennings 1964). Such observations<br />

may suggest that japonica rices be grown in the tropics. In fact, ponlai nces,<br />

japonica rices developed in Taiwan and adapted to subtropical climates, perform<br />

well in tropical regions. However, the direct introduction <strong>of</strong> japonica rices into the<br />

tropics was not favored for several reasons. First, most japonica varieties mature<br />

very early because <strong>of</strong> daylength and temperature sensitivity, resulting in stunted<br />

growth and low yield. Only ponlai rices are adapted to tropical climates; consequently,<br />

the amount <strong>of</strong> genetic resources is quite limited. Second, people in<br />

tropical Asia prefer the cooking quality <strong>of</strong> indica rices. Third, japonica rices lack<br />

the grain dormancy desirable for tropical varieties that mature toward the end <strong>of</strong><br />

the monsoon season.<br />

With this consideration, it was logical to make hybrids between japonicas and<br />

indicas to transfer the good characters <strong>of</strong> japonica rices to indica rices. In the<br />

1950s, the Food and Agriculture Organization (FAO) organized an indicajaponica<br />

hybridization program with its headquarters at the Central <strong>Rice</strong> Research<br />

Institute (CRRI) in Cuttack, India (Parthasarathy 1972). From this program came<br />

ADT27 in India and Mahsuri and Malinja in Malaysia. However, the over-all<br />

results made little impact on rice improvement programs in South and Southeast<br />

Asia.<br />

The warm tropical climate was considered another factor responsible for low<br />

yields. Because high temperatures increase respiratory losses, net dry matter<br />

production — as a balance between photosynthesis and respiration — was thought<br />

to be inevitably lower under higher temperatures. Low rice yields in southwestern<br />

Japan were attributed to warm climates. For example, yields were correlated with<br />

solar radiation and air temperature by the following formula (Murata 1964):<br />

Y = S [1.20 – 0.021 ( t – 21.5) 2 ],<br />

(6.1)<br />

where Y is grain yield (kg/10 a [ a = 0.01 ha]), and S and t are solar radiation<br />

(cal/cm 2 per day) and daily mean temperature (°C), in August and September. The<br />

equation indicates that temperatures higher or lower than 21.5°C will decrease

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