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Yoshida - 1981 - Fundamentals of Rice Crop Science

Yoshida - 1981 - Fundamentals of Rice Crop Science

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PHYSIOLOGICAL ANALYSIS OF RICE YIELD 247<br />

The water table <strong>of</strong> the field used in this experiment was 80 cm below the soil<br />

surface in winter and 18 cm in summer. The rice grain yield is usually around 4.5<br />

t/ha.<br />

The researchers found that a low number <strong>of</strong> sunshine-hours during ripening<br />

resulted in a low percentage <strong>of</strong> filled spikelets. To overcome this problem, they<br />

shifted the crop season 1 month earlier. Component technologies included drainage,<br />

deep plowing, and application <strong>of</strong> 19–56 t compost/ha (Table 7.6). AS a<br />

consequence, applied nitrogen exceeded 400 kg/ha. There was not much difference<br />

in grain yield between the treatments when the field was poorly drained.<br />

When the field received adequate drainage, grain yield increased from 6.6 to 10.0<br />

t/ha.<br />

Another example <strong>of</strong> high yielding technology is the intensive use <strong>of</strong> topdressing.<br />

Nitrogen topdressing is normally done at panicle initiation or flag-leaf stage<br />

in Hokkaido, Japan. Shiga and Miyazaki (1977) studied the nitrogen absorption<br />

process <strong>of</strong> a high yielding crop and simulated that process by topdressing. Nitrogen<br />

topdressing was given 5 times between panicle initiation and heading, and<br />

the amount <strong>of</strong> nitrogen applied each time was increased stepwise from 5 kg N/ha at<br />

panicle initiation to 10 kg N/ha 5 days later, to 15 kg N/ha after another 5 days after<br />

that, to 30 kg N/ha at flag-leaf stage (15 days after panicle initiation), and 30 kg<br />

N/ha at heading. By this technique, grain yield was increased from 3.8 t/ha without<br />

fertilizer nitrogen to 9.7 t/ha with a simulated topdressing. The simulated topdressing<br />

produced high yields <strong>of</strong> more than 8.8 t/ha over 5 years.<br />

7.6. FACTORS LlMlTlNG THE PRESENT YIELD PLATEAU<br />

Improving grain-filling is one way to increase grain yield. The filled-spikelet<br />

percentage is about 85%, even under favorable conditions, which means that grain<br />

yield might be increased by 15%. Of the 15% unfilled spikelets, however, around<br />

5–10% are unfertilized, and difficult to eliminate. An increase in photosynthesis<br />

(source activity) relative to spikelet number (sink size) would probably increase<br />

filled-spikelet percentage by only 5–10% and, hence, yield by a similar degree.<br />

Thus, the yield to be achieved by improving gain-filling would be much lower<br />

than the estimated potential yield.<br />

It is possible to increase yield by increasing the number <strong>of</strong> spikelets per unit <strong>of</strong><br />

land area. In other words, it is important to increase yield capacity when yield<br />

capacity is defined as the product <strong>of</strong> spikelet number and potential weight per<br />

grain. To examine this possibility, it must be known whether grain filling, as<br />

determined by the amount <strong>of</strong> solar radiation during ripening, is already limiting to<br />

yield. The question implies that the number <strong>of</strong> spikelets per unit <strong>of</strong> land area may<br />

be increased through improved cultural practices or by genetic manipulation, but a<br />

considerable portion <strong>of</strong> the spikelets produced may remain unfilled because <strong>of</strong><br />

limited solar radiation.<br />

In Japan. the percentage <strong>of</strong> ripened grains decreases when the number <strong>of</strong><br />

spikelets per square meter is increased (Matsushima 1970, Wada 1959). Thus,

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