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Yoshida - 1981 - Fundamentals of Rice Crop Science

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216 FUNDAMENTALS OF RICE CROP SCIENCE<br />

considered the first high yielding indica rice variety. It demonstrated that increasing<br />

the yield potential <strong>of</strong> indica rices can be achieved by improvements within<br />

indica rices. Thus, it pointed the way for tropical rice breeding.<br />

In 1962, IRRI began research at Los Baños, Philippines. Its plant breeders<br />

crossed Dee-geo-woo-gen and Peta at IRRI. Peta, originating in Indonesia and<br />

popularly grown in the Philippines, is a tall, high-tillering variety. From the cross<br />

IR8 was selected and released in 1966. IR8 is erect leaved, high tillering,<br />

photoperiod insensitive, and about 100 cm tall with stiff culms. It responds well to<br />

nitrogen and produces about 6 t/ha in the wet season and 9 t/ha in the dry.<br />

Occasionally, the yield exceeds 10 t/ha. Figure 6.1 compares the nitrogen<br />

response <strong>of</strong> IR8 and traditional varieties. IR8 is considered the first high yielding<br />

indica rice variety adapted to tropical climates. With good management dwarf<br />

rices such as IR8 have enormous potential to boost rice yields in Asia. Their impact<br />

was elegantly described as Dwarf rice — a giant in tropical Asia (Chandler 1968).<br />

Dee-geo-woo-gen was used as a genetic source <strong>of</strong> semidwarf stature in both<br />

TN1 and IR8. Its usefulness is comparable to that <strong>of</strong> Norin 10 for Mexican wheat.<br />

Both Dee-geo-woo-gen and Norin 10 originated in Asia.<br />

6.3. PLANT-TYPE CONCEPT<br />

6.3.1. Evolution <strong>of</strong> concept<br />

The term plant type refers to a set <strong>of</strong> morphological characters associated with the<br />

yielding ability <strong>of</strong> rice varieties. The plant-type concept in rice, similar to the wheat<br />

ideotype (Donald 1968), emerged gradually from many observations and comparisons<br />

between low and high yielding varieties in Japan (Baba 1954, 1961;<br />

Tsunoda 1964).<br />

Better varieties <strong>of</strong> crop species must have been selected for higher yields in the<br />

past. In practice, the high yields have been achieved by increased inputs, particularly<br />

nitrogen fertilizers. Hence, varieties must have been selected on the basis <strong>of</strong><br />

their response to nitrogen fertilizers. Tsunoda (1964) compared low and high<br />

yielding varieties <strong>of</strong> rice, sweet potato, and soybean, and summarized their<br />

morphological characteristics:<br />

• Low nitrogen responders have long, broad, thin, drooping, pale-green<br />

leaves, and tall, weak stems.<br />

• High nitrogen responders have erect, short, narrow, thick, dark-green leaves,<br />

and short, sturdy stems.<br />

Tsunoda postulated from physiological knowledge <strong>of</strong> crop photosynthesis that<br />

the thick, dark-green leaves lose less light through reflection. Reduced leaf size<br />

and erect habit permit uniform light distribution to all leaves and reduce respiration.<br />

Consequently, dry matter and yield increase even under low light conditions.<br />

The uniform light penetrations and the short, sturdy culms minimize lodging; thus<br />

there is little or no yield loss once panicles are well developed (Tsunoda 1964,<br />

Jennings 1964).

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