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Yoshida - 1981 - Fundamentals of Rice Crop Science

Yoshida - 1981 - Fundamentals of Rice Crop Science

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GROWTH AND DEVELOPMENT OF THE RICE PLANT 47<br />

1.39. The relationship between<br />

temperature and days to heading<br />

for 4 photoperid-insensitive varieties<br />

grown in temperaturecontrolled<br />

glasshouses (IRRI<br />

1975).<br />

become commercially available and are being used for many such studies.<br />

The responses <strong>of</strong> four photoperiod-insensitive varieties to constant temperatures<br />

are shown in Figure 1.39. There are clear varietal differences in the number <strong>of</strong> days<br />

to heading for each temperature in terms <strong>of</strong> temperature summation or heat<br />

requirement. Yukara, a variety from northern Japan, has a much lower temperature<br />

summation value than IR8 and IR26, both tropical varieties. The shape <strong>of</strong> the<br />

response curves is basically the same and typical for many other crops. Within a<br />

daily mean temperature range <strong>of</strong> 21°-30°C. the number <strong>of</strong> days to heading is not<br />

linearly related to temperature. When temperature drops from 24° to 21°C, there is<br />

a sharp increase in days to heading. For example, the number <strong>of</strong> days to heading<br />

for IR26 increases from 96 days at 24°C to 134 days at 21°C. In other words, a<br />

temperature drop by 1°C causes a 13-day delay in heading. When the temperature<br />

is increased above 24°C, however, days to heading decreases to 91 days at 27°C<br />

and to 86 days at 30°C. In other words, a temperature rise <strong>of</strong> 1°C above 24°C<br />

shortens the number <strong>of</strong> days to heading by less than 2 days, These results appear to<br />

indicate that temperatures above 24°C are much less effective than those below<br />

24°C, suggesting the existence <strong>of</strong> a ceiling temperature. This idling effect <strong>of</strong> high<br />

temperature on growth duration is similar to that observed in the field. When<br />

IR747B2-6 is grown in the field throughout the year at Los Baños, Philippines,<br />

where monthly mean temperatures change from 25.3°C in January to 29.4°C in<br />

May, it matures in 95 days regardless <strong>of</strong> planting time.<br />

A generalized relationship between temperature and length <strong>of</strong> time required to<br />

complete development is shown in Figure 1.40. The curve indicates a temperature<br />

below which the plant will not grow. Also, there is an intermediate optimum<br />

temperature that permits the most rapid development. Above the optimum,

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