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Yoshida - 1981 - Fundamentals of Rice Crop Science

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236 FUNDAMENTALS OF RICE CROP SCIENCE<br />

7.3. Relationship between total number <strong>of</strong> spikelets per square meter. grain yield. filled-spikelet<br />

percentage, and grain weight <strong>of</strong> variety IR8 (<strong>Yoshida</strong> et al 1972).<br />

number per square meter was the most important component limiting yield in this<br />

experiment.<br />

A similar conclusion was obtained from IR8 crops grown at Los Baños (Fig.<br />

7.3). The grain yield increased linearly with increasing spikelet number per square<br />

meter while filled-spikelet percentage and 1,000-grain weight remained fairly<br />

constant regardless <strong>of</strong> spikelet number. At some locations and in some weather<br />

conditions, filled-spikelet percentage may be more limiting to yield than spikelet<br />

number per square meter (<strong>Yoshida</strong> and Parao 1976). The 1,000-grain weight may<br />

affect yield to some extent but rarely becomes limiting under most conditions.<br />

Therefore, under given conditions it is important to examine the causes <strong>of</strong> yield<br />

variation in components, particularly spikelet number per square meter and filledspikelet<br />

percentage.<br />

7.3.2. Spikelet number per unit land area<br />

Spikelet number per unit land area is affected by:<br />

• Cultural practices — spacing (seeding density) and nitrogen application.<br />

• Growth characteristics — tillering (seedling emergence).<br />

• Climatic conditions — solar radiation and temperature.<br />

Under subnormal cultural practices, the spikelet number per unit land area can<br />

be increased by increasing plant density and nitrogen application. Optimum<br />

spacing varies with tillering capacity and growth duration <strong>of</strong> a variety. A higher<br />

plant density is required for low-tillering, early-maturing varieties, and low<br />

nitrogen levels. The relationships between spikelet number per unit land area and<br />

amount <strong>of</strong> nitrogen absorbed by the crop have been discussed in Chapter 2 (Fig.<br />

2.9).

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