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Yoshida - 1981 - Fundamentals of Rice Crop Science

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MINERAL NUTRITION OF RICE 141<br />

3.12. Carbohydrate content <strong>of</strong> leaf sheath plus culm, nitrogen content <strong>of</strong> active leaf blade,<br />

and active leaf blade ratio <strong>of</strong> IR8, 1967 dry season (<strong>Yoshida</strong> and Ahn 1968).<br />

photosynthetic production increases relative to the absorption rate <strong>of</strong> ammonia, the<br />

nitrogen content decreases and surplus photosynthates accumulate.<br />

The close and inverse correlation between nitrogen nutrition and carbohydrate<br />

accumulation suggests that the latter can be used as an indication <strong>of</strong> nitrogen<br />

deficiency. A starch iodine test has been proposed to determine the need for<br />

nitrogen topdressing at panicle initiation (Kiuchi 1968).<br />

3.7.5. Partial productive efficiency<br />

One <strong>of</strong> the central themes in plant nutrition is to find the particular growth stage<br />

when supplying a nutrient produces the maximum yield per unit amount <strong>of</strong> nutrient<br />

absorbed.<br />

Partial productive efficiency <strong>of</strong> nitrogen for grain production is defined as:<br />

(3.21)<br />

where Y ( n ) = grain weight <strong>of</strong> the treatment<br />

corresponding to period ( n ),<br />

Y ( n – 1) = grain weight <strong>of</strong> the treatment<br />

corresponding to period ( n – 1 ),<br />

N ( n ) = total nitrogen uptake <strong>of</strong> the treatment<br />

corresponding to period ( n ), and<br />

N ( n – 1) = total nitrogen uptake <strong>of</strong> the treatment<br />

corresponding to period ( n – 1).<br />

In other words, partial productive efficiency is the amount <strong>of</strong> grain produced per<br />

unit amount <strong>of</strong> nitrogen absorbed at a particular growth stage. Thus, the greater the<br />

value <strong>of</strong> partial productive efficiency, the higher the efficiency <strong>of</strong> nitrogen<br />

absorbed to produce grains. Partial productive efficiency <strong>of</strong> nitrogen for straw is<br />

also defined in the same way as for grain.<br />

The solution culture technique provides a convenient means to study partial<br />

productive efficiency. In solution culture, the nutrient supply can be controlled

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